STATUS OF BOCACCIO OFF CALIFORNIA IN 2007

Alec D. MacCall

National Oceanic and Atmospheric Administration

National Marine Fisheries Service
Southwest Fisheries Science Center
Fishery Ecology Division
110 Shaffer Rd.
Santa Cruz, CA 95060
<[email protected]>

May, 2008

Table of Contents
2 Executive Summary
14 Assessment
36 Appendix A. Responses to STAR Panel requests
44 Appendix B. Responses to comments and recommendations in 2003 and 2005 STAR
reports
52 Appendix C. Data file for model STATc2007
62 Appendix D. Parameter file for model STATc2007
66 Appendix E. Summary tables of model results
Executive Summary – Bocaccio

Approach: This assessment was conducted primarily as an “update” which follows the
methodology and assumptions of the 2003 bocaccio assessment as closely as possible. The main
differences from the previous assessment are addition or revision of recent data, and revision of
the historical commercial catches. One additional model is added, based on a prior distribution
of stock-recruitment steepness. The assessment used the original Stock Synthesis model (SS1),
and does not develop an equivalent new Stock Synthesis 2 (SS2) version of the assessment.
Accordingly, some features of SS2 output, such as precision estimates, do not appear in this
assessment.

Stock: Bocaccio rockfish (Sebastes paucispinis) occurring in waters off the state of California.
For management purposes, the stock may be considered to reside in U.S. waters south of Cape
Mendocino. This stock assessment treats the resource in Southern and Central California as a
combined unit.

Catches: Catches have declined steeply from the 1970s, reflecting both a long-term decline in
abundance and progressive restrictions on harvest of bocaccio (Table ES1, Figure ES1). Values
of catches since 2000 are imprecise because of management-induced discarding. Recent
discards in the trawl fishery have been monitored; for lack of better information, discard rates in
other commercial fisheries are assumed to be similar those for the trawl fishery. Discards in the
recreational fishery were obtained from RecFIN. Details are given in Table ES2.

Data and assessment: This assessment follows the methodology and assumptions of the 2003
bocaccio assessment as closely as possible. This assessment uses the original Stock Synthesis
model (SS1, synl32r.exe, compiled 4/2/2003), and does not develop an equivalent new Stock
Synthesis 2 (SS2) version of the assessment.

Input data extend back to 1951. Data include catches from five fisheries segments
reflecting three statewide commercial gears (trawl, setnet, hook&line), and separate southern
California and central/northern California recreational fisheries, length compositions from six
sources (all five fisheries segments, and the Triennial Survey), and six indexes of abundance
(trawl logbook CPUE, three recreational CPUEs, Triennial Survey abundance, and CalCOFI
larval index of spawning output). The assumed natural mortality rate (M) was 0.15/yr in
accordance with the 2003 assessment.

Unresolved problems and major uncertainties: Within the scope of this update assessment,
there were no unresolved problems or uncertainties. The STATc model developed in the 2003
assessment is the focus of the update, with more limited consideration of the STARb1 and
STARb2 models. Differences among the three models are described in Table ES3. One
additional model developed in this document is based on the maximum posterior density
estimate of bocaccio stock-recruitment steepness (h = 0.44) from a meta-analysis of rockfish
stock

2
Table ES1. Summary of historical bocaccio catches (mtons, including discards)
Trawl Hook&Line Setnet RecSOUTH RecNORTH Total
1950 1287 200 0 39 86 1612
1960 2163 351 0 63 125 2702
1970 1660 298 0 289 204 2451
1980 3641 335 216 1755 178 6037
1990 1144 497 793 233 91 2451
1995 377 69 281 44 3 777
1996 288 93 92 67 26 573
1997 230 58 35 49 107 480
1998 73 42 39 29 23 209
1999 45 21 7 71 53 197
2000 54 21 2 52 60 189
2001 59 35 4 60 49 207
2002 41 7 0 76 8 132
2003 1 2 0 11 0 14
2004 11 9 0.3 59 2 82
2005 23 26 0.1 32 6 87
2006 5 20 0.2 31 11 67

Table ES2. Estimated recent fishery removals (mtons) of bocaccio. Parentheses indicate value used in 2003 assessment.
TRAWL H&L SETNET RecSouth RecCen Total
Retained Disc Discard** Est Retained Est Retained Est A + B1 A + B1 Est
Rate*
2000 20.1 34.0 54(54) 7.0 21(21) 0.7 2(2) 52 60 189(187)
2001 13.7 45.7 59(37) 7.8 35(23) 0.9 4(2) 60 49 207(187)
2002 18.2 56% 41(99) 3.0 7(17) 0.2 0(1) 76 8 132(201)
2003 0.2 79% 1 0.5 2 0.0 0 11 0 14
2004 6.2 42% 11 5.5 9 0.3 0.3 60 2 83
2005 3.9 83% 23 4.4 26 0.1 0.1 32 6 87
2006 0.9 83% 5 3.4 20 0.2 0.2 31 11 68
* Discard rate from J. Hastie (email 2007); **Discards from J. Hastie (email 2005); 2006 discard rate assumed same as 2005

3
Table ES3. Summary of 2003 bocaccio models. Bold type indicates updated aspects of the
models.

M = 0.15
Years: background, 1950 to 2002
Recruitments (age 1):
STAR B1: expval 1951-59, individual 1960-2001, expectval 2002, 2003; SRR lambda=0
STAR B2: expval 1951-69, individual 1970-2001, expectval 2002, 2003; SRR lambda=0
STAT C: expval 1951-59, individual 1960-2001, expectval 2002, 2003: SRR lambda=0.1
Age bins: 1 to 21+
Length bins: 24, 26, …… 66, 68, 72,76, 80+
Growth: Von Bertalanffy fitted in model, separate male and female curves
Length CVs: 0.107 at age 1.5, 0.033 at age 99

Modeled Segments: Selectivity form First LF Last LF Nyears Sexes Used?

Trawl Dbl. Logistic 1978 2002 25 yes all
Hook and Line Dbl. Logistic 1980 2002 22 yes all
Set Net Dbl. Logistic 1978 2002 18 yes all
Recreational South Dbl. Logistic 1975 2002 24 no all
Recreational North Dbl. Logistic 1980 2002 23 no all
Triennial Trawl Survey Dbl. Logistic 1986 2001 6 yes not in STAR B1

Abundance Indexes Selectivity source First Last Nyears CV Used?

RecFIN CPUE North Rec North 1980 2002 20 0.67 not in STAR B2
CDFG CPUE North Rec North 1987 1998 12 0.37 all
RecFIN CPUE South Rec South 1980 2002 20 0.71 not in STAR B2
Trawl CPUE (north) Trawl 1982 1996 15 0.32 all
Triennial Trawl Triennial 1977 2001 9 0.81 not in STAR B1
CalCOFI Larval Spawn Ogive 1951 2003 47 0.68 all

Recruitment Indexes Selectivity source First Last Nyears CV Used?

Power Plant Ent’nment age 1 1972 2000 29 2.10 no
Cen Cal Juvenile Trawl age 1 1983 2002 20 2.05 no
Rec Pier CPUE age 1 1980 2002 20 3.29 no

assessments (Dorn, pers. comm. 2007), with full emphasis (lambda = 1) on the Beverton-Holt
model fit to the recruitment estimates, with all other aspects of the model similar to the STATc
model.

Reference points: Values in this discussion are from the STATc model; values for all four
models are given in Table ES4. Population reproductive potential is measured as spawning
output (units of billion eggs). Unfished abundance cannot be estimated reliably from historical
stock and recruitment due to lack of curvature in the relationship. An imprecise estimate of
unfished spawning output was obtained by multiplying the average age-1 recruitment (1951 to
1986) by unfished SPR, giving 13572 billion eggs.

4
 The 50%SPR exploitation rate (Catch/Biomass age 1+) is 0.0630, which is used as a
proxy Fmsy rate by the PFMC. Proxy Bmsy (40% of Bunfished) corresponds to an approximate
equilibrium total biomass of 31341 mtons, and if this is fished at proxy Fmsy, the MSY is
estimated to be 1974 mtons. Although calculations related to MSY are imprecise, estimates vary
relatively little among alternative models and methods of calculation. The overfishing threshold
is Fmsy, in this case the proxy value of F(SPR50%), and corresponding catch levels are the ABC
values given in the rebuilding projections.

Stock biomass: The estimated history of the biomass (age 1+) and spawning output (billion
eggs) estimated by the four alternative models are shown in Figures ES2 and ES3 and values are
given in Table ES5. Notably the three models (STARb1, STATc and STATcMPDh) that are
allowed to estimate early biomass indicate that biomass was near the minimum stock size
threshold at the beginning of the assessment period, ca. 1950. This is not surprising, given that
the assumed pre-1950 “background” catch was 2000 mtons, which slightly exceeds estimated
MSY. CalCOFI larval abundances indicate a relatively low biomass in the 1950s, and a
substantial increase in spawning abundance during the 1960s.

Recruitment: The estimated history of recruitment (omitting earlier years with little data to
inform estimates) is shown in Figure ES4 and values are given in Table ES6. The strong 1999
year class was followed by a moderately strong 2003 year class. Strength of the 2004 and 2005
year classes is not estimated from data, but rather is taken from the stock-recruitment
relationship. The recruitment values for these years are not substantiated by other sources of
information, and these values may be overly optimistic.

Exploitation status: The history of exploitation rates is shown in Figure ES5 and ES6, and
values are given in Table ES6. From the STATc model, the estimated spawning output in 2006
is 1727 billion eggs, or 12.7% of the estimated unfished level. The estimated 2006 total biomass
(age 1+) is 10752 mtons. The 2006 exploitation rate of 0.0062 was far below the reference
exploitation rate of 0.0630 that is the maximum fishing mortality threshold under the SPR50%
proxy (see Figure ES1). At the Fmsy proxy, the STATc model gives a 2006 catch (ABC) of 677
mtons (this is also the overfishing threshold) and a “40-10" policy optimum yield (OY) of 193
mtons.

Management performance: The 2006 OY was set at 218 mtons, the retained catch was about
42 mtons, and the estimated total catch including discards was 68 mtons (Tables ES2 and ES7).
Including mortality of estimated discards, estimated total kill in 2006 was 67 mtons. Thus,
recent management has been achieving total removals well below target levels, and far below
maximum levels. A ten-year history of management performance is given in Table ES7.

5
Table ES4. Management reference points for bocaccio.
Model
units STARb1 STARb2 STATc STATcMPDh
Steepness(h) ca. 0.2 ca. 0.2 ca. 0.2 0.44

Unfished Reference Points

Reference source Ravg51-86 Ravg51-86 Ravg51-86 SRR
Spawning Output billion eggs 13563 13132 13572 12591
Summary (1+) Biomass mtons 71104 68894 71195 66036
Mean Recruitment at age 1 thousands 5451 5270 5449 5039
SPR(F=0) 2.488 2.492 2.491 2.502

Current status (2006)

Spawning Output billion eggs 2075 1430 1727 2049
Spawning Output at SB40% billion eggs 5425 5253 5429 5036
Relative depletion 15.3% 10.9% 12.7% 16.3%
Summary (1+) Biomass mtons 14559 9582 10752 13661
2006 Catch (including discards) mtons 67 67 67 67
Exploitation rate 0.0046 0.0070 0.0062 0.0049
Overfishing threshold (ExpRate at SPR50%) 0.0633 0.0631 0.0630 0.0633
ABC mtons 922 605 677 865
OY (40-1010) mtons 426 66 193 401

Reference Points based on SPR50% proxy at SB40%

Spawning Output at SB40 billion eggs 5425 5253 5429 5036
SPRmsy proxy 50% 50% 50% 50%
Exploitation rate at SPR50% 0.0633 0.0631 0.0630 0.0633
Approx Bsummary at SB40% mtons 30928 30313 31341 29146
given Exploitation rate at SPR50%
Yield with SPR50% at SB40% mtons 1958 1913 1974 1845

Reference Points based on estimated MSY values from SRR

Smsr, Spawning Output at MSY billion eggs undefined undefined undefined 4549
Smsr/Sunfished 36%
R at Smsr thousands 4138
SPRmsy 1.0992
rel SPRmsy 44%
Exploitation rate at SPRmsy 0.0768
Bsummary at Smsy mtons 29671
MSY mtons undefined undefined undefined 2279

Forecasts: The first year of projection was 2006, so that the recruitment of age 1 fish in 2007
and later was obtained by random resampling of R/S value from the spawning years of 1969
through 2003 (Figure ES7, Table ES8). Catches were fixed at the observed 67 mtons in 2006,
and at a projected 151 mtons in both 2007 and 2008. Beginning in 2009, the projections use a
constant fishing rate corresponding to an SPR of 77.7% (2009 rebuilding OY would be 288
mtons), without reversion to the 40-10 harvest policy upon reaching the rebuilding target of B40.
Based on 2000 simulations, approximately half of the projections reach the rebuilding target by
2023, and 67% of the simulations were rebuilt by the current statutory rebuilding target date of
2026. If the probability of attaining the rebuilding target by 2026 is reduced to 50%, the SPR
could be decreased to 66.4%, allowing larger catches (in which case, 2009 rebuilding OY would
be 468 mtons).

Decision tables: No table was developed because this stock is subject to a rebuilding plan.

6
Table ES5. History of bocaccio biomass and spawning output.

Total biomass (age 1+) Spawning output (billion eggs)

STARb1 STARb2 STATc STATcMPDh STARb1 STARb2 STATc STATcMPDh
mtons mtons mtons mtons sp out rel depl sp out rel depl sp out rel depl sp out rel depl
avg 71104 68894 71195 70069 13563 100% 13132 100% 13572 100% 13368 100%
unexpl
1950 23644 39688 22625 23029 3764 28% 6871 52% 3580 26% 3650 27%
1960 16575 33291 16405 16320 2546 19% 5666 43% 2359 17% 2429 18%
1970 44285 32447 43288 42458 8306 61% 5099 39% 7910 58% 7450 56%
1975 30504 28662 30969 30069 4980 37% 4115 31% 5034 37% 4829 36%
1980 29064 29148 29561 28792 3467 26% 3406 26% 3600 27% 3426 26%
1985 13229 13434 13504 13091 2160 16% 2239 17% 2256 17% 2136 16%
1990 8994 8876 9039 8880 1140 8% 1170 9% 1179 9% 1117 8%
1995 5510 4842 5348 5474 820 6% 757 6% 820 6% 812 6%
1996 5257 4466 5037 5223 819 6% 730 6% 808 6% 812 6%
1997 5222 4297 4944 5198 826 6% 712 5% 804 6% 820 6%
1998 5136 4091 4796 5117 835 6% 697 5% 802 6% 830 6%
1999 5276 4126 4888 5274 881 6% 718 5% 836 6% 877 7%
2000 6562 4918 5882 6511 928 7% 739 6% 871 6% 925 7%
2001 7412 5416 6522 7374 970 7% 757 6% 901 7% 970 7%
2002 8611 6137 7422 8559 1050 8% 797 6% 958 7% 1050 8%
2003 9712 6788 8213 9634 1277 9% 938 7% 1134 8% 1274 10%
2004 11341 7720 9283 11047 1599 12% 1145 9% 1386 10% 1592 12%
2005 12805 8563 10024 12275 1863 14% 1309 10% 1585 12% 1852 14%
2006 14559 9582 10752 13661 2075 15% 1430 11% 1727 13% 2049 15%

7
Table ES6. Recent trends in recruitment (thousands) and exploitation rate.
Recruitment at age 1 Exploitation rate (C/B1+)
Year STARb1 STARb2 STATc STATcMPDh STARb1 STARb2 STATc STATcMPDh
1995 879 690 796 867 14.0% 16.0% 14.5% 14.1%
1996 509 369 435 497 10.8% 12.7% 11.2% 10.8%
1997 1061 843 1006 1109 9.2% 11.1% 9.7% 9.2%
1998 334 205 245 317 4.0% 5.0% 4.3% 4.0%
1999 384 297 368 432 3.7% 4.8% 4.0% 3.7%
2000 7385 4977 5944 7043 2.9% 3.8% 3.2% 2.9%
2001 56 50 50 249 2.8% 3.8% 3.2% 2.8%
2002 625 442 481 607 1.5% 2.2% 1.8% 1.5%
2003 861 469 489 710 0.1% 0.2% 0.2% 0.1%
2004 4602 2433 2732 3480 0.7% 1.1% 0.9% 0.7%
2005 2651 1907 917 2279 0.7% 1.0% 0.9% 0.7%
2006 3080 2176 1049 2524 0.5% 0.7% 0.6% 0.5%
* Recruitment values for 2005 and 2006 are expected values from SRR, not estimated.

Table ES7. Recent history of management performance.

Commercial Recreational Total ABC OY
Year Catch Discard Total Catch Discard Total Catch Discard Total
1997 323 * 323 145 11 156 468 11 479 265 265
1998 154 * 154 52 0 52 206 0 206 230 230
1999 73 * 73 120 4 124 193 4 197 230 230
2000 28 49 77 103 9 112 128 58 189 164 100
2001 22 76 98 103 6 109 125 82 207 122 100
2002 21 27 48 82 2 84 103 32 132 122 100
2003 1 2 3 9 2 11 10 4 14 244 <20
2004 12 8 20 55 8 62 66 18 82 400 199
2005 8 41 49 34 4 38 42 45 87 566 307
2006 5 20 25 37 5 42 42 25 67 549 306
2007 53** 98** 151** 602 218
2008 618 218
* Discarded commercial catch was not estimated and is assumed to be negligible.
** Projected as of August, 2007 (John. DeVore, pers. comm.)

Research and data needs: The recommendations presented here are from the STAT Team;
STAR Panel reports also contain recommendations on this subject. Future bocaccio assessments
should utilize the Stock Synthesis 2 model, and time-varying growth rates should be explored
because of apparent interdecadal variability in progressions of modal lengths. A two-area model
(north and south of Pt. Conception) may help distinguish the state of the resource in those two
areas, but migration patterns and rates are not known well enough to project rebuilding
trajectories separately for the two areas. The southern California segment may be a vitally
important source of migrants to central California waters. The CalCOFI larval survey, including
central California stations, is critical to future bocaccio assessments. Tracking intra-annual
patterns of gonadal states could improve its interpretation and eventually lead to calibrated
estimates of true abundance. The STAT recommends against pursuing trawl-based abundance
estimates, due to poor ability to sample rocky habitats preferred by bocaccio. An acoustic-
optical system being developed at the SWFSC may be suitable for estimating bocaccio
abundance.

8
Table ES8. Median projected abundances of bocaccio, at F(SPR=77.7%) beginning in 2009, without reversion to 40-10 policy.
Estimates are based on model STATc and future projections do not include imprecision in estimated 2006 status. Bold values indicate
rebuilt status. Catch is observed value in 2006, and is assumed in 2007-08.

Year SPR Catch ABC Depletion Spawning Output

projected median median 5% median 95% 5% median 95%
2006 0.939 67 677 12.7% 12.7% 12.7% 1727 1727 1727
2007 0.871 151 693 13.8% 13.8% 13.8% 1872 1872 1873
2008 0.823 218 704 14.9% 14.9% 14.9% 2015 2016 2024
2009 0.777 288 718 15.6% 15.7% 16.4% 2117 2128 2221
2010 0.777 302 753 15.9% 16.3% 18.9% 2156 2209 2564
2011 0.777 323 806 15.9% 17.0% 22.1% 2154 2298 2994
2012 0.777 354 882 15.8% 17.8% 25.7% 2137 2419 3480
2013 0.777 387 964 15.6% 18.9% 28.8% 2115 2561 3900
2014 0.777 426 1062 15.6% 20.2% 32.5% 2109 2744 4404
2015 0.777 467 1165 15.6% 21.8% 35.8% 2113 2960 4856
2016 0.777 507 1263 15.7% 23.7% 40.2% 2130 3212 5452
2017 0.777 546 1361 16.1% 25.9% 44.5% 2183 3505 6036
2018 0.777 586 1460 16.4% 27.9% 49.2% 2226 3782 6665
2019 0.777 622 1550 16.9% 30.1% 54.0% 2291 4079 7320
2020 0.777 661 1649 17.4% 32.4% 59.7% 2365 4396 8092
2021 0.777 723 1804 18.1% 34.5% 66.6% 2454 4680 9033
2022 0.777 772 1926 18.7% 37.1% 74.4% 2532 5025 10084
2023 0.777 826 2060 19.3% 39.9% 83.3% 2622 5408 11295
2024 0.777 890 2220 20.2% 43.0% 91.2% 2743 5829 12367
2025 0.777 936 2334 20.9% 46.4% 101.2% 2839 6285 13721
2026 0.777 1018 2538 21.9% 49.4% 110.4% 2962 6699 14961

9
 9000
8000 Foreign
7000
Recreational
Catch (mtons)

6000
Commercial
5000
4000
3000
2000
1000
0
50 55 60 65 70 75 80 85 90 95 100 105
Year

Figure ES1. History of bocaccio catches, showing foreign, recreational and

commercial components. A catch of 2000 mtons is assumed prior to 1950.

60000
Total Biomass (age 1+, mtons)

STATc
50000
STATcMPDh
40000 STARb1
STARb2
30000

20000

10000

0
1950 1960 1970 1980 1990 2000 2010
Year

Figure ES2. History of total biomass (age 1+) of bocaccio estimated by four
alternative models.

10
 9000
8000 STATc
Spawning Output (billion eggs)

7000 STATcMPDh
6000 STARb1
5000 STARb2

4000
3000
2000
1000
0
1950 1960 1970 1980 1990 2000 2010
Year

Figure ES3. History of spawning output of bocaccio estimated by four alternative

models.

30000
STATc
Spawning Output (billion eggs)

25000
STATcMPDh
20000 STARb1
STARb2
15000

10000

5000

0
1968
1970
1972
1974
1976
1978
1980
1982
1984
1986
1988
1990
1992
1994
1996
1998
2000
2002
2004
2006

Year

Figure ES4. Trends in bocaccio recruitment (thousand fish at age 1) estimated by four
alternative models. Indicated year is age 1, spawning occurred in previous year.

11
 0.35
STATc
0.3 STATcMPDh
Exploitation Rate (C/Bage 1+)

0.25 STARb1
STARb2
0.2

0.15

0.1

0.05

0
1950 1960 1970 1980 1990 2000 2010
Year

Figure ES5. Trends in bocaccio exploitation rate indicated by four alternative

models.

5
ExpRate/ExpRate(SPR50%)

0
0 0.5 1 1.5
SpOut/SpOut40%

Figure ES6. Historical exploitation rates and spawning outputs relative to target
values. Plot begins in 1951 (near center) and ends in 2006 (lower left). Model is
STATc.

12
 spawning Output (billion eggs) 10000

8000

6000

4000

2000

0
1980 1990 2000 2010 2020 2030
Year

Figure ES7. Projected spawning output under a constant SPR=77.7% rebuilding

policy. Trajectories are 5, 25, 50 (median, bold line), 75 and 95 percentiles; broken
line is B40 rebuilding target.

13
 Status of bocaccio off California in 2007

Introduction

A full stock assessment of bocaccio off California was last conducted in 2003 (MacCall
2003a), and an update assessment was conducted in 2005 (MacCall 2005a). Although initial
plans were to convert the model to SS2 and do an extensive bocaccio assessment in 2007, a
variety of unanticipated events resulted in a revision of the work plan. Rather than cancelling
the assessment altogether, an assessment equivalent to an update in scope was proposed for 2007
(email MacCall to Clarke, 10/18/06). However, the STAR Panel review schedule was not
modified.

This assessment is equivalent to an “update” assessment, but the STAR Panel review
allows additional leeway to consider alternative models. According to the TOR for stock
assessment updates, an update stock assessment must carry forward its fundamental structure
from a model that was previously reviewed and endorsed by a STAR Panel. Accordingly, this
assessment adheres to the model frameworks established in 2003, and is similar to the update
developed in 2005 (MacCall 2005a). With regard to the specific requirements for an update
assessment, there must be similarity in:
a) the particular sources of data used,
b) the analytical methods used to summarize data prior to input to the model,
c) the software used in programming the assessment,
d) the assumptions and structure of the population dynamics model underlying the stock
assessment,
e) the statistical framework for fitting the model to the data and determining goodness of
fit,
f) the procedure for weighting the various data components, and
g) the analytical treatment of model outputs in determining management reference points,
including Fmsy, Bmsy, and B0.
The present assessment satisfies all of these requirements.

For clarity of presentation, this document treats current results relative to the original
2003 assessment, without attempting to reconcile very minor changes from the 2005 update,
most of which were due to use of incomplete data from 2004 and 2005 in the latter assessment.
Specifications of the three models developed in the 2003 assessment are summarized in Table 1.
The STARb1 and STARb2 models omit portions of the data. The STATc model includes all of
the data sources and is the accepted basis of management decisions. One additional model
described later in this document is a variant of the STATc model where the stock-recruitment
curve has increased influence on the model.

14
 Fishery data

Catches

Revision of commercial catches: The time series of commercial catches was revised, due
to recovery of misplaced data from landings sampling during the late-1970's to early 1990's. The
new (2007) catch values are compared with the previous (2003) catch values in Figure 1.
Annual values of catches by individual fishery segments are given in the Stock Synthesis data
file (Appendix B).

Historical catches by major fishery segments are shown in Figure 2, and detail of
landings and discard in recent years are given in Table 2. Five distinct fishery segments are
recognized in this assessment: commercial trawl, hook and line and set net gears, and
recreational fisheries north and south of Pt. Conception. Recent estimates of recreational catch
and discards (Table 2) were obtained from the RecFIN database. Commercial catches were
obtained from the CALCOM database (Don Pearson, SWFSC, pers. comm.), and estimated rates
of bycatch/discard from the trawl fishery were provided by Jim Hastie (NWFSC, pers. comm.).
Because gears other than trawl were not observed sufficiently, the trawl discard rate was applied
to all commercial gears. The estimated three-year 2000-2002 cumulative catch and discard (528
mtons) is slightly lower than in the 2003 assessment (575 mtons). Annual catch rates declined
in the subsequent 2003-2006 period due to severe restrictions imposed by management.

Length Compositions

Recreational fisheries: Length compositions of retained bocaccio are available from the
southern California recreational fishery in 2003-2006 (Figure 3), and in the central California
recreational fishery in 2004-2006 (Figure 4). Sample sizes are given in Table 3. In central
California, the strong 1999 year class remains dominant through 2006. In southern California,
the 1999 year class declines in importance after 2004, and the 2003 year class dominates the size
composition in 2005-2006.

Commercial fisheries: The severe decline in commercial landings of bocaccio has

resulted in few length composition samples of commercially-caught bocaccio (Figure 5, Table
3). Useful data exist only from the trawl fishery in 2004 and the hook and line fishery in 2006.
Samples exist for the set net fishery in 2004 and hook and line fishery in 2005 but were too small
to be used.

Fishery-Dependent Abundance Indexes

No attempt was made to update the recreational fishery CPUE abundance indexes
because of difficulty interpreting catch rates under the strong restrictions that were placed on
landing bocaccio.

15
 Fishery-Independent Data: Surveys and Indexes

Triennial Survey

The information from the Triennial Survey is unchanged from the 2005 update, but is
included here for completeness. A Triennial Trawl Survey was conducted in 2004 (data
provided by Mark Wilkins, AFSC, and Beth Horness, NWFSC, pers. comm.). The length
composition of bocaccio taken in that survey is show in Figure 5. As was done previously, I
used a simple log-transformed GLM to produce year-specific indexes of abundance. This
approach allows a consistent interpretation of the survey results even though the Conception area
was not sampled in 1980, 1983 and 1986. The GLM predicts stratum means with fixed area,
depth and year effects (Figure 6), and a minor error for year 1980 was corrected as in the 2005
update. The new index values are consistent with those used in the 2003 assessment.

CalCOFI Survey

The 2003 assessment included the January 2003 CalCOFI ichthyoplankton survey, but
did not include the April 2003 survey. This assessment includes both CalCOFI surveys through
April 2006 (Richard Charter, SWFSC, pers. comm.). Annual sample sizes for the southern and
central California portions of the survey are given in Table 2. As before, a delta-lognormal
GLM with fixed year, month and station effects was use to produce annual index values (Figure
7). Consistency between values in the 2003 assessment and this assessment is shown in Figure
8. The index value for 2003 decreased slightly when the April survey data for that year were
included.

Recruitment Indexes

In its review of the 2003 assessment, the STAR Panel recommended excluding use of
recruitment indexes. Those indexes are not used in this assessment, and updated values were not
calculated.

Assessment Model

The assessment was conducted using the “Stock Synthesis 1” length-based maximum
likelihood model (synl32r.exe, compiled 4/2/2003), and is directly comparable to the 2003
assessment. As in the 2003 assessment, natural mortality rate is set at M=0.15. All three of the
models developed in the 2003 assessment (STARB1, STARB2 and STATC; see Table 1 for
model details) are updated here.

This assessment includes consideration of one additional model, based on an assumed

value of Mace-Doonan steepness (h) in a Beverton-Holt stock-recruitment relationship (SRR).
Martin Dorn (ms in prep) recently conducted a Bayesian meta-analysis of Beverton-Holt SRRs
for a number of west coast rockfish stocks. The bocaccio information in that analysis was based

16
on the STATc model used for the 2005 update. Dorn provided a maximum posterior density
(mpd) estimate of h = 0.44 for bocaccio (Dorn email 6/4/07). That value of steepness is used
here in model STATcMPDh. The only differences from the STATc model are the assumed
value of h = 0.44 and an increase in emphasis (lambda) on the stock-recruitment residuals to 1.

Model Results

Model results, including a retrospective view of results for 2003 from the new model, are
compared in Table 4, and more details are given in Tables 5 and 6. Abundance trajectories,
recruitments and exploitation rates are shown in Figures 9-13. Fits to abundance time series are
shown in Figure 14 and fits to recent length compositions are shown in Figure 15. All four
models are in general agreement for the most recent 30 years (models STARb1 and STARb2
differ in specification of early recruitments).

Reference points: Values in this discussion are from the STATc model; values for all four
models are given in Tables 4 and 5. Population reproductive potential is measured as spawning
output (units of billion eggs). Except for model STATcMPDh with its explicit stock-recruitment
relationship (SRR), unfished abundance cannot be estimated reliably from historical stock and
recruitment due to lack of curvature in the estimated relationship. An imprecise estimate of
unfished spawning output was obtained by multiplying the average age-1 recruitment (1951 to
1986) by unfished SPR, giving 13572 billion eggs. The SRR from model STATcMPDh is used
to estimate the values of reference points for that model (Table 5). Estimated values of reference
points vary little among alternative models.

The 50%SPR exploitation rate (Catch/Biomass age 1+) is 0.0630, which is used as a
proxy Fmsy rate by the PFMC. Proxy Bmsy (40% of Bunfished) corresponds to an approximate
equilibrium total biomass of 31341 mtons, and if this is fished at proxy Fmsy, the MSY is
estimated to be 1974 mtons. Although calculations related to MSY are imprecise, estimates vary
relatively little among alternative models and methods of calculation. The overfishing threshold
is Fmsy, in this case the proxy value of F(SPR50%), and corresponding catch levels are the ABC
values given in the rebuilding projections.

Stock biomass: The estimated history of the biomass (age 1+) and spawning output
(billion eggs) estimated by the four alternative models are shown in Figures 9 and 10 and recent
values are given in Table 6. Notably the three models (STARb1, STATc and STATcMPDh) that
are allowed to estimate early biomass indicate that biomass was near the minimum stock size
threshold at the beginning of the assessment period, ca. 1950. This is not surprising, given that
the assumed pre-1950 “background” catch was 2000 mtons, which slightly exceeds estimated
MSY. CalCOFI larval abundances indicate a relatively low biomass in the 1950s, and a
substantial increase in spawning abundance during the 1960s. Abundance has approximately
doubled since rebuilding began in year 2000.

Recruitment: The estimated history of recruitment (omitting earlier years with little data
to inform estimates) is shown in Figure 11 and values are given in Table 6. The strong 1999

17
year class was followed by a moderately strong 2003 year class. Strength of the 2004 and 2005
year classes is not estimated from data, but rather is taken from the stock-recruitment
relationship. The recruitment values for these years are not substantiated by other sources of
information, and these values may be overly optimistic.

Exploitation status: The history of exploitation rates is shown in Figure 12 and 13, and
values are given in Table 6. From the STATc model, the estimated spawning output in 2006 is
1727 billion eggs, or 12.7% of the estimated unfished level. The estimated 2006 total biomass
(age 1+) is 10752 mtons. The 2006 exploitation rate of 0.0062 was far below the reference
exploitation rate of 0.0630 that is the maximum fishing mortality threshold under the SPR50%
proxy (see Figure 13). At the Fmsy proxy, the STATc model gives a 2006 catch (ABC) of 677
mtons (this is also the overfishing threshold) and a “40-10" policy OY of 193 mtons.

Retrospective patterns: Retrospective patterns given in Table 4 indicate that 2003

estimates from model STARb2 are nearly unchanged in the 2007 assessment (estimated 2003
spawning output increases by 2%). The 2007 STATc model results for 2003 provide upward
revisions of 2003 abundance estimates, with an 11% increase in estimated 2003 spawning
output. The 2007 STARb1 model indicates substantial upward revision of its 2003 estimates,
with a 44% upward revision of estimated 2003 spawning output. For a more detailed year-by-
year analysis of retrospective patterns, the reader is referred to the 2005 assessment document.

Rebuilding Projections

Projections used the SSC Default Rebuilding Analysis (Version 2.10b) programmed by
Andre Punt (program available at http://www.fish.washington.edu/people/punt/software.html).
The first year of projection was 2006, so that the recruitment of age 1 fish in 2007 and later was
obtained by random resampling of R/S value from the spawning years of 1969 through 2003
(Figure 16, Table 7). Catches were fixed at the observed 67 mtons in 2006, and at a projected
151 mtons in both 2007 and 2008. Beginning in 2009, the projections use a constant fishing rate
corresponding to an SPR of 77.7% (2009 rebuilding OY would be 288 mtons), without reversion
to the 40-10 harvest policy upon reaching the rebuilding target of B40. Based on 2000
simulations, approximately half of the projections reach the rebuilding target by 2023, and 67%
of the simulations were rebuilt by the current statutory rebuilding target date of 2026. If the
probability of attaining the rebuilding target by 2026 is reduced to 50%, the SPR could be
decreased to 66.4%, allowing larger catches (in which case, 2009 rebuilding OY would be 468
mtons).

18
 Research and Data Needs

The recommendations presented here are from the STAT Team; STAR Panel reports also
contain recommendations on this subject. Future bocaccio assessments should utilize the Stock
Synthesis 2 model, and time-varying growth rates should be explored. Although a two-area
model (north and south of Pt. Conception) is worth exploring to distinguish the state of the
resource in those two areas, migration patterns and rates are not known well enough to project
rebuilding trajectories separately for the two areas. The southern California segment may prove
to be less depleted, but may be a vitally important source of migrants to central California
waters. Continuation of the CalCOFI larval survey, including central California stations, is
critical to future bocaccio assessments. Tracking intra-annual patterns of gonadal states could
improve its interpretation and eventually lead to calibrated estimates of true abundance. The
STAT recommends against pursuing trawl-based abundance estimates, due to poor ability to
sample rocky habitats preferred by bocaccio. An acoustic-optical survey system being
developed at the SWFSC in La Jolla may be suitable for estimating bocaccio abundance.

Acknowledgements

I would like to thank people who provided updated data sets used in this assessment: Don
Pearson, Richard Charter, Beth Horness, Mark Wilkins, and Jim Hastie. My thanks also extend
to the much larger number of people who did the field work and data processing that underlie the
summary information I received.

References

MacCall, A. 2003a. Status of bocaccio off California in 2003. Pacific Fishery Management
Council. <http://www.pcouncil.org/groundfish/gfsafe0803/bocaccio.pdf>

MacCall, A. 2003b. Bocaccio rebuilding analysis for 2003. Pacific Fishery Management
Council. <http://www.pcouncil.org/groundfish/gfsafe0803/bocaccioreb.pdf>

MacCall, A. 2005a. Status of bocaccio off California in 2005. Pacific Fishery Management
Council. <http://www.pcouncil.org/groundfish/gfsafe0406/bocaccio2005final.pdf>

MacCall, A. 2005b. Bocaccio rebuilding analysis for 2005. Pacific Fishery Management
Council. <http://www.pcouncil.org/groundfish/gfsafe0406/BOCrebuild2005.pdf>

19
Table 1. Summary of 2003 bocaccio models. Bold type indicates updated aspects of the models.

M = 0.15
Years: background, 1950 to 2002
Recruitments (age 1):
STAR B1: expval 1951-59, individual 1960-2001, expectval 2002, 2003; SRR lambda=0
STAR B2: expval 1951-69, individual 1970-2001, expectval 2002, 2003; SRR lambda=0
STAT C: expval 1951-59, individual 1960-2001, expectval 2002, 2003: SRR lambda=0.1
Age bins: 1 to 21+
Length bins: 24, 26, …… 66, 68, 72,76, 80+
Growth: Von Bertalanffy fitted in model, separate male and female curves
Length CVs: 0.107 at age 1.5, 0.033 at age 99, interpolated on mean length at age

Modeled Segments: Selectivity form First LF Last LF Nyears Sexes Used?

Trawl Dbl. Logistic 1978 2002 25 yes all
Hook and Line Dbl. Logistic 1980 2002 22 yes all
Set Net Dbl. Logistic 1978 2002 18 yes all
Recreational South Dbl. Logistic 1975 2002 24 no all
Recreational North Dbl. Logistic 1980 2002 23 no all
Triennial Trawl Survey Dbl. Logistic 1986 2001 6 yes not in STAR B1

Abundance Indexes Selectivity source First Last Nyears CV* Used?

RecFIN CPUE North Rec North 1980 2002 20 0.67 not in STAR B2
CDFG CPUE North Rec North 1987 1998 12 0.37 all
RecFIN CPUE South Rec South 1980 2002 20 0.71 not in STAR B2
Trawl CPUE (north) Trawl 1982 1996 15 0.32 all
Triennial Trawl Triennial 1977 2001 9 0.81 not in STAR B1
CalCOFI Larval Spawn Ogive 1951 2003 47 0.68 all

Recruitment Indexes Selectivity source First Last Nyears CV Used?

Power Plant Ent’nment age 1 1972 2000 29 2.10 no
Cen Cal Juvenile Trawl age 1 1983 2002 20 2.05 no
Rec Pier CPUE age 1 1980 2002 20 3.29 no

* Re-tuning of CVs is automatic based on RMSE of fit (log scale).

20
Table 2. Estimated recent fishery removals (mtons) of bocaccio. Parentheses indicate value used in 2003 assessment.
TRAWL H&L SETNET RecSouth RecCen Total
Retained Disc Discard** Est Retained Est Retained Est A + B1 A + B1 Est
Rate*
2000 20.1 34.0 54(54) 7.0 21(21) 0.7 2(2) 52 60 189(187)
2001 13.7 45.7 59(37) 7.8 35(23) 0.9 4(2) 60 49 207(187)
2002 18.2 56% 41(99) 3.0 7(17) 0.2 0(1) 76 8 132(201)
2003 0.2 79% 1 0.5 2 0.0 0 11 0 14
2004 6.2 42% 11 5.5 9 0.3 0.3 60 2 83
2005 3.9 83% 23 4.4 26 0.1 0.1 32 6 87
2006 0.9 83% 5 3.4 20 0.2 0.2 31 11 68
* Discard rate from J. Hastie (email 2007); **Discards from J. Hastie (email 2005); 2006 discard rate assumed same as 2005

Table 3. Sample size and model tuning information for updated information sources. Starred samples were small and not used.
Length Compositions Year Nobs Neff Units Nsamples
Trawl 2004 110 78 Fish 9
Set Net 2004 17* n.a. Fish 2
Hook&Line 2005 11* n.a. Fish 2
2006 78 13 Fish 10
So Calif Recreational 2003 122 84 Fish
2004 889 86 Fish
2005 571 85 Fish
2006 1330 88 Fish
Cen Calif Recreational 2004 80 14 Fish
2005 73 13 Fish
2006 49 9 Fish

Triennial Survey 2004 33 23 Hauls

Abundance Index Year Cen Calif So Calif Units

CalCOFI 2003 52 92 Stations
2004 49 88 Stations
2005 86 191 Stations
2006 40 148 Stations

21
Table 4. Comparison of model results.

Model Total Biomass Spawning Spawn Output Spawn Output

mt, age 1+ Output Unfished rel to Unfished

STAR B1 (exclude Triennial Survey index)

2003 original 8913 1136 13412 8.5%
2003 new 9712 1277 9.4%
2007 new 14559 2075 13563 15.3%

STAR B2 (exclude Recreational CPUE)

2003 original 5455 733 13064 5.6%
2003 new 6783 938 7.1%
2007 new 9582 1430 13132 10.9%

STAT C (use all abundance indexes)

2003 original 7133 984 13387 7.4%
2003 new 8213 1134 8.4%
2007 new 10752 1727 13572 12.7%

STAT C MPDh (use all abundance indexes)

2003 original
2003 new 9634 1274 9.5%
2007 new 13661 2049 13368 15.3%

22
Table 5. Management reference points for bocaccio.
Model
units STARb1 STARb2 STATc STATcMPDh
Steepness(h) ca. 0.2 ca. 0.2 ca. 0.2 0.44

Unfished Reference Points

Reference source Ravg51-86 Ravg51-86 Ravg51-86 SRR
Spawning Output billion eggs 13563 13132 13572 12591
Summary (1+) Biomass mtons 71104 68894 71195 66036
Mean Recruitment at age 1 thousands 5451 5270 5449 5039
SPR(F=0) 2.488 2.492 2.491 2.502

Current status (2006)

Spawning Output billion eggs 2075 1430 1727 2049
Spawning Output at SB40% billion eggs 5425 5253 5429 5036
Relative depletion 15.3% 10.9% 12.7% 16.3%
Summary (1+) Biomass mtons 14559 9582 10752 13661
2006 Catch (including discards) mtons 67 67 67 67
Exploitation rate 0.0046 0.0070 0.0062 0.0049
Overfishing threshold (ExpRate at SPR50%) 0.0633 0.0631 0.0630 0.0633
ABC mtons 922 605 677 865
OY (40-1010) mtons 426 66 193 401

Reference Points based on SB40%

Spawning Output at SB40% billion eggs 5425 5253 5429 5036
Recruitment at SB40 thousands 4322
SPR resulting in SB40% 47%
Exploitation rate at SB40% 0.0707
Bsummary at R(SB40%) mtons 29565
Yield with SPR(SB40%) at SB40% undefined undefined undefined 2090

Reference Points based on SPR50%

Spawning Output at SPR50% billion eggs 5668
Recruitment at SB(SPR50%) thousands 4531
SPRmsy proxy 50% 50% 50% 50%
Exploitation rate at SPR50% 0.0633 0.0631 0.0630 0.0633
Bsummary at R(SB(SPR50%)) mtons 32803
Yield with SPR50% at SB(SPR50%) undefined undefined undefined 2076

Reference Points based on SPR50% proxy at SB40%

Spawning Output at SB40 billion eggs 5425 5253 5429 5036
SPRmsy proxy 50% 50% 50% 50%
Exploitation rate at SPR50% 0.0633 0.0631 0.0630 0.0633
Approx Bsummary at SB40% mtons 30928 30313 31341 29146
given Exploitation rate at SPR50%
Yield with SPR50% at SB40% mtons 1958 1913 1974 1845

Reference Points based on estimated MSY values from SRR

Smsr, Spawning Output at MSY billion eggs undefined undefined undefined 4549
Smsr/Sunfished 36%
R at Smsr thousands 4138
SPRmsy 1.0992
rel SPRmsy 44%
Exploitation rate at SPRmsy 0.0768
Bsummary at Smsy mtons 29671
MSY mtons undefined undefined undefined 2279

23
Table 6. Results of model STATc for recent years. Approximate values at MSY assume
F(SPR=0.5) at B=0.4Bunfished where Bunfished is estimated from the unfished biomass
resulting from average recruitment from 1951 to 1986.
Year Spawning Relative Total age1+ Recruits Catch Exploitation
Output Abundance Biomass at age 1 Rate
avg value unfished 13572 100% 71230 5456 0 0
approx value at MSY 5429 40% 31340 2182 1974 6.30%
1995 820 6.0% 5348 796 774 14.5%
1996 808 6.0% 5037 435 566 11.2%
1997 804 5.9% 4944 1006 479 9.7%
1998 802 5.9% 4796 245 206 4.3%
1999 836 6.2% 4888 368 197 4.0%
2000 871 6.4% 5882 5944 189 3.2%
2001 901 6.6% 6522 50 207 3.2%
2002 958 7.1% 7422 481 132 1.8%
2003 1134 8.4% 8213 489 14 0.2%
2004 1386 10.2% 9283 2732 82 0.9%
2005 1585 11.7% 10024 917 87 0.9%
2006 1727 12.7% 10752 1049 67 0.6%

Table 7. Median projected abundances of bocaccio, at SPR of 77.7% beginning in 2009, without
reversion to 40-10 policy upon attainment of rebuilding target. Estimates are based on model
STATc and future projections do not include imprecision in estimated 2006 status. Bold values
indicate rebuilt status. Catch is observed value in 2006, and is assumed in 2007-08.
Year SPR Catch ABC Depletion Spawning Output
projected median median 5% median 95% 5% median 95%
2006 0.939 67 677 12.7% 12.7% 12.7% 1727 1727 1727
2007 0.871 151 693 13.8% 13.8% 13.8% 1872 1872 1873
2008 0.823 218 704 14.9% 14.9% 14.9% 2015 2016 2024
2009 0.777 288 718 15.6% 15.7% 16.4% 2117 2128 2221
2010 0.777 302 753 15.9% 16.3% 18.9% 2156 2209 2564
2011 0.777 323 806 15.9% 17.0% 22.1% 2154 2298 2994
2012 0.777 354 882 15.8% 17.8% 25.7% 2137 2419 3480
2013 0.777 387 964 15.6% 18.9% 28.8% 2115 2561 3900
2014 0.777 426 1062 15.6% 20.2% 32.5% 2109 2744 4404
2015 0.777 467 1165 15.6% 21.8% 35.8% 2113 2960 4856
2016 0.777 507 1263 15.7% 23.7% 40.2% 2130 3212 5452
2017 0.777 546 1361 16.1% 25.9% 44.5% 2183 3505 6036
2018 0.777 586 1460 16.4% 27.9% 49.2% 2226 3782 6665
2019 0.777 622 1550 16.9% 30.1% 54.0% 2291 4079 7320
2020 0.777 661 1649 17.4% 32.4% 59.7% 2365 4396 8092
2021 0.777 723 1804 18.1% 34.5% 66.6% 2454 4680 9033
2022 0.777 772 1926 18.7% 37.1% 74.4% 2532 5025 10084
2023 0.777 826 2060 19.3% 39.9% 83.3% 2622 5408 11295
2024 0.777 890 2220 20.2% 43.0% 91.2% 2743 5829 12367
2025 0.777 936 2334 20.9% 46.4% 101.2% 2839 6285 13721
2026 0.777 1018 2538 21.9% 49.4% 110.4% 2962 6699 14961

24
 all Commercial gears Hook & Line

6000 1200
old values (2003) old values (2003)
5000
Landings (mtons)

1000

Landings (mtons)
revised (2007) revised (2007)
4000 800
3000 600
2000 400
1000 200
0 0
75 80 85 90 95 100 105 75 80 85 90 95 100 105
Year Year

Set Net Trawl

800 5000
700 old values (2003) old values (2003)
Landings (mtons)

Landings (mtons)
revised (2007) 4000 revised (2007)
600
500 3000
400
300 2000
200
1000
100
0 0
75 80 85 90 95 100 105 75 80 85 90 95 100 105
Year Year

Figure 1. Comparison of revised commercial landings of bocaccio with values used in the 2003 assessment.

25
 9000
8000 Foreign
7000
Recreational
Catch (mtons)

6000
Commercial
5000
4000
3000
2000
1000
0
50 55 60 65 70 75 80 85 90 95 100 105
Year

Figure 2. History of bocaccio catches, showing foreign, recreational and

commercial components. A catch of 2000 mtons is assumed prior to 1950.

2006
2005
2004
2003
0.4 2002
2001
0.2
2000
0
24
30
36
42
48
54
60
66
76

Length (FL-cm)

Figure 3. Length composition of bocaccio landed by the southern California

recreational fishery.

26
 2006
2005
2004
2003
0.4 2002
2001
0.2
2000
0
24
30
36
42
48
54
60
66
76
Length (FL-cm)

Figure 4. Length composition of bocaccio landed by the central California recreational

fishery.

2006 H&L
0.3
2004 Trawl
0.2

0.1 2004 Triennial

0
24

28

32

36

40

44

48

52

56

60

64

68

76

Length (FL-cm)

Figure 5. Other length compositions used in the 2007 bocaccio update.

27
 8
7
6
Ln Triennial Index

5
4
3 2005 GLM
2 2003 GLM

1
0
1975 1980 1985 1990 1995 2000 2005 2010
Year

Figure 6. Triennial Trawl Survey index of bocaccio abundance (note log scale).

3.5

2.5
CalCOFI Index

1.5

0.5

0
1940 1950 1960 1970 1980 1990 2000 2010
Year

Figure 7. CalCOFI Survey index of bocaccio spawning output. Error bars are ± 1SE.

28
 Index rel to 1951-1994 avg 1.6
1.4
2007 assmt
1.2 2003 assmt
1
0.8
0.6
0.4
0.2
0
1985 1990 1995 2000 2005 2010
year

Figure 8. Comparison of updated recent CalCOFI Index values with values used in
the 2003 bocaccio assessment (standardized for comparison). The 2003 point was
based on incomplete data available in the 2003 assessment.

60000
Total Biomass (age 1+, mtons)

STATc
50000
STATcMPDh
40000 STARb1
STARb2
30000

20000

10000

0
1950 1960 1970 1980 1990 2000 2010
Year

Figure 9. History of total biomass (age 1+) of bocaccio estimated by four alternative
models.

29
 9000
8000 STATc
Spawning Output (billion eggs)

7000 STATcMPDh
6000 STARb1
5000 STARb2

4000
3000
2000
1000
0
1950 1960 1970 1980 1990 2000 2010
Year

Figure 10. History of spawning output of bocaccio estimated by four alternative

models.

30000
STATc
Spawning Output (billion eggs)

25000
STATcMPDh
20000 STARb1
STARb2
15000

10000

5000

0
1968
1970
1972
1974
1976
1978
1980
1982
1984
1986
1988
1990
1992
1994
1996
1998
2000
2002
2004
2006

Year

Figure 11. Trends in bocaccio recruitment (thousand fish at age 1) estimated by four alternative
models. Indicated year is age 1, spawning occurred in previous year.
30
 0.35
STATc
0.3 STATcMPDh
Exploitation Rate (C/Bage 1+)

0.25 STARb1
STARb2
0.2

0.15

0.1

0.05

0
1950 1960 1970 1980 1990 2000 2010
Year

Figure 12. Trends in bocaccio exploitation rate indicated by four alternative

models.

5
ExpRate/ExpRate(SPR50%)

0
0 0.5 1 1.5
SpOut/SpOut40%

Figure 13. Historical exploitation rates and spawning outputs relative to target
values. Plot begins in 1951 (near center) and ends in 2006 (lower left). Model is
STATc.

31
 RecFIN CPUE North RecFIN CPUE South

3 4.5
4
2.5
3.5
2 3
CPUE

CPUE
2.5
1.5
2
1 1.5
1
0.5
0.5
0 0
75 80 85 90 95 100 105 75 80 85 90 95 100 105
Year Year

CDF&G Rec CPUE North Trawl Logbook CPUE

7 180
6 160
140
5
120
4
CPUE

CPUE
100
3 80
60
2
40
1 20
0 0
75 80 85 90 95 100 105 75 80 85 90 95 100 105
Year Year

Triennial Trawl Survey CalCOFI Survey

1400 3
Larval Abundance Index

1200 2.5
Swept Area Index

1000
2
800
1.5
600
1
400
200 0.5

0 0
75 80 85 90 95 100 105 50 60 70 80 90 100 110
Year Year

Figure 13. Model STATc2007 fits to abundance indexes and surveys. Triennial trawl survey in
2004 and CalCOFI indexes for 2003-2006 are the only new data since 2003.

32
 Recreational South 2003

0.25

0.2
frequency

0.15
0.1
0.05
0
24

28

32

36

40

44

48

52

56

60

64

68

76
Length Bin

Recreational South 2004 Recreational North 2004

0.16 0.16
0.14 0.14
0.12 0.12

frequency
frequency

0.1 0.1
0.08 0.08
0.06 0.06
0.04 0.04
0.02 0.02
0 0

24

28

32

36

40

44

48

52

56

60

64

68

76
24

28

32

36

40

44

48

52

56

60

64

68

76

Length Bin Length Bin

Recreational South 2005 Recreational North 2005

0.2 0.25
0.2
0.15
frequency

frequency

0.15
0.1
0.1
0.05 0.05
0 0
24

28

32

36

40

44

48

52

56

60

64

68

76

24

28

32

36

40

44

48

52

56

60

64

68

76
Length Bin Length Bin

Recreational South 2006 Recreational North 2006

0.25 0.2
0.2
0.15
frequency

frequency

0.15
0.1
0.1
0.05 0.05

0 0
24

28

32

36

40

44

48

52

56

60

64

68

76

24

28

32

36

40

44

48

52

56

60

64

68

76

Length Bin Length Bin

Figure 14. STATc model fits to recreational fishery length compositions collected since 2003.

33
 Trawl 2004 Hook & Line 2006

0.3 0.14

0.25 0.12
0.1
0.2
frequency

frequency
0.08
0.15
0.06
0.1
0.04
0.05 0.02
0 0
24

28

32

36

40

44

48

52

56

60

64

68

76

24

28

32

36

40

44

48

52

56

60

64

68

76
Length Bin Length Bin

Triennial Trawl (females) 2004 Triennial Trawl (males) 2004

0.12 0.12

0.1 0.1

0.08 0.08
frequency

frequency
0.06 0.06

0.04 0.04

0.02 0.02

0 0
24

28

32

36

40

44

48

52

56

60

64

68

76

24

28

32

36

40

44

48

52

56

60

64

68

76
Length Bin Length Bin

Figure 15. STATc model fits to commercial fishery and Triennial Trawl Survey length compositions.

34
 spawning Output (billion eggs) 10000

8000

6000

4000

2000

0
1980 1990 2000 2010 2020 2030
Year

Figure 16. Projected spawning output under a constant SPR=77.7% rebuilding

policy. Trajectories are 5, 25, 50 (median, bold line), 75 and 95 percentiles; broken
line is B40 rebuilding target.

35
Appendix A. Responses to STAR Panel requests

Some requests are repeats of earlier requests, but were given a new sequence number:
The response to Request B is given as the responses to requests H and L.
The response to Request E is given as the response to request J.
The response to Request F is given as the response to request I.
The response to Request G is given as the response to request K.

STAR Panel request A. Re-examine the historical rockfish catch (back to 1916) using the
ratio of bocaccio to total rockfish to estimate historical catches.

Reason: This will determine if the assumed equilibrium catch of 2000t is reasonable.

STAT response: The assumed 2000 ton historical catch was based on the few years prior to
1950. The entire history of rockfish catch in California is shown in Figure A1. Species
compositions from early years indicate that bocaccio comprised approximately 50% of the total
rockfish. The equilibrium value should be based on a weighted average catch, with recent years
receiving more weight than much earlier years. Using an F=M assumption, an exponentially
declining relative weight of exp(-0.3*yrs) gives a weighted pre-1951 average rockfish catch of
3150 mtons, or about 1575 mtons of bocaccio, which is somewhat below the value assumed in
the assessment.

7000
Calif total rockfish (mtons)

6000

5000
4000

3000
2000

1000
0
1915

1926

1930

1940

1945

1950

1955
1020

1935

Year

Figure A1. Rockfish catches from early California landings

records. Horizontal line is bocaccio catch assumed in the
model.

36
STAR Panel request C. Do a model run using steepness=0.44, lambda=1 on the S/R curve and
remove the recent recreational CPUE values. Retune the CPUE and survey S.E.

Reason: This is a working model to be used in the subsequent runs. Removing the recent cpue
values removes the conflict with the triennial survey.

STAT response: This model run (termed the STAR model) was done as requested, and results
are shown in Figure A2. The effect of removing recent recreational CPUE values has little
effect on the model. Assuming steepness h=0.44 with full emphasis on the SRR has an effect
mainly on pre-1970 estimates of abundance, including estimated unfished abundance. The
STAR model gives an estimated unfished spawning output of 9763 and a current spawning
output of 1761, for a currrent depletion level of 18%. The STATc model (h=0.2) does not give
a SRR-based estimate of unfished spawning output, but the standard SSC procedure of applying
unfished SPR to the average early recruitments gives an unfished spawning output of 13572,
and the current spawning output of 1660 is a depletion level of 12%.

The STAT considers the Triennial Survey, which is conducted in inappropriate habitat for
bocaccio, to be the least reliable source of abundance information in the bocaccio model, and
disagrees with the STAR Panel in the rationale for downweighting the recent recreational CPUE
values. Adoption of a steepness of h=0.44 is arbitrary, and is one of a wide range of plausible
values rom Dorn’s meta-analysis. Proper treatment of the uncertainty in steepness requires a
full MCMC integration of model outcomes, but cannot be done in SS1.

9000 STATc2007
8000 STAR (del recent Rec CPUE)
7000
Spawning Output

STAR, w/ SRR
6000
5000
4000
3000
2000
1000
0
45 55 65 75 85 95 105
Year

Figure A2. Model requested by STAR Panel request C,

showing cumulative effect of deleting recent CPUE, and
then assuming steepness h=0.44.

37
STAR Panel request D. Do sensitivity runs to equilibrium catches of 1000 and 3000t using the
working model (Request C). Output the biomass trajectory and depletion.

Reason: Determine the effect of the magnitude of equilibrium catch on estimated abundance.

STAT response: The model values of spawning output are shown in Figure A3. Alternative
assumptions of early catches has an effect mainly on pre-1970 estimates of abundance,
including estimated unfished abundance. Depletion and related reference points are as follows:

Model Unfished Sp Output Current Sp Output Relative Depletion

STAR, eq C = 1000 9376 1971 21%
STAR, eq C = 2000 9763 1761 18%
STAR, eq C = 3000 11654 1682 14%

9000
8000 STAR, eq C = 3000
7000 STAR, eq C = 1000
Spawning Output

6000
STAR, eq C = 2000
5000
4000
3000
2000
1000
0
45 50 55 60 65 70 75 80 85 90 95 100 105 110
Year

Figure A3. STAR model with alternative assumed values of

pre-1951 catch.

STAR Panel request E. Do a new model run starting in 1916 with no equilibrium catch and
ramping up catch to 2000t in 1930. This will be done using the working model (Request C).
Output the biomass trajectories and depletion.

Reason: To demonstrate the effect of assuming equilibrium conditions as opposed to assuming

the population was not in equilibrium conditions with a similar magnitude of catch.

STAT response: Array limitations in SS1 do not allow the model to start as early as 1916. The
STAR Panel produced a modified request J below.

38
STAR Panel request H. (Repeats request B) Determine the co-occurrence of other species in
RecFIN trips that caught bocaccio from 1993-2006. For trips that caught bocaccio, produce the
proportion of those trips that also caught other species (only the top 20 or so).

Reason: Attempt to distinguish near shore from offshore fishing using co-occurrence of species
in the RecFIN source, to confirm no change in recreational fishery selection patterns in recent
years.

STAT response: (Southern California is given under Request L). The data were divided into
four time periods, the 1980s, the 1990, 2000-2002, when bocaccio catches were restricted but
fishing areas were unrestricted, and 2004-2006 when fishing was restricted mostly to nearshore
areas (no samples were taken in 2003). As expected, the species co-occurrences tend to show a
higher probability of catching nearshore species in the latter time period. However, the STAT
disagrees with the STAR Panel interpretation. Any increase in selectivity of nearshore fish in
central California is irrelevant to the estimated strength of the 2003 year class because it was
not seen in central California length comps–it was only seen in southern California.

In the following table, a BOLD entry indicates a value higher than in any preceding column,
and a small italic entry indicates a value smaller than in any preceding column.
species\years 1980's 1990's 2000-02 2004-06
n bocaccio trips (Cen CA) 241 238 29 92
yellowtail rockfish 0.618 0.895 0.828 0.587
blue rockfish 0.324 0.647 0.310 0.924
lingcod 0.390 0.622 0.310 0.826
canary rockfish 0.494 0.655 0.621 0.228
vermilion rockfish 0.232 0.517 0.379 0.870
rosy rockfish 0.373 0.706 0.587 0.511
starry rockfish 0.340 0.643 0.483 0.739
olive rockfish 0.199 0.521 0.276 0.826
copper rockfish 0.154 0.382 0.310 0.815
gopher rockfish 0.058 0.193 0.103 0.457
widow rockfish 0.261 0.529 0.448 0.261
China rockfish 0.071 0.126 0.103 0.435
black rockfish 0.033 0.034 0.087
greenspotted rockfish 0.402 0.361 0.621 0.098
brown rockfish 0.041 0.076 0.034 0.272
yelloweye rockfish 0.299 0.214 0.276 0.033
chub (Pacific) mackerel 0.116 0.239 0.141
Pacific sanddab 0.083 0.214 0.138 0.185
greenstriped rockfish 0.336 0.248 0.345 0.011
chilipepper 0.432 0.155 0.379 0.054
kelp greenling 0.017 0.013 0.152
speckled rockfish 0.104 0.277 0.207 0.076
cabezon 0.021 0.017 0.185
flag rockfish 0.075 0.176 0.241 0.228

39
STAR Panel request I. (Repeats request F) Do a new model assuming a logistic selectivity
pattern (estimated) for the triennial survey and using the working model (Request C). Produce
a table of likelihoods for all components, biomass trajectories and depletion. Show the fits to
the triennial proportion at length data, CPUE series, and surveys.

Reason: To see the effect of the current selectivity pattern on the estimates of recruitment in the
recent period.

STAT response: The resulting “logistic” selectivity is uniform for all sizes, as there is no
ascending limb. The log-likelihood for the Triennial Survey length comps is 11.7 units worse,
and the overall model is 11.2 units worse. This selectivity curve would be rejected by
conventional goodness-of-fit criteria. The table of likelihood values is given below.

Model
Source Type base logistic difference
TWL Length Comps -539.144 -538.437 0.707
H&L Length Comps -203.397 -203.141 0.256
SetNet Length Comps -258.783 -258.286 0.497
RECLso Length Comps -319.271 -320.059 -0.788
RECLnor Length Comps -267.985 -268.389 -0.404
TRIENNIAL Length Comps -83.199 -94.901 -11.702
RecFINno CPUE (N) 1.003 0.939 -0.064
NoCalDFG CPUE (N) 8.198 8.183 -0.015
RecFINso CPUE (N) -3.867 -3.885 -0.018
TrawlLogbooks CPUE (B) 9.036 9.345 0.309
TRIENNIAL SURVEY (BIOMASS) -5.772 -5.549 0.223
CALCOFI SURVEY (SSB) -1.722 -1.582 0.14
BEV-HOLT SRR MODEL -25.411 -25.787 -0.376
TOTAL -1690.32 -1701.55 -11.2329

40
STAR Panel request J. (a modification of request E). Do a new model run starting in 1930
(due to SS1 constraints) with no equilibrium catch and catches after 1930 until the start of the
measured catches set at 2000t. This will be done using the working model (Request C). Output
the biomass trajectories and depletion.

Reason: To demonstrate the effect of assuming equilibrium conditions as opposed to assuming

the population was not in equilibrium conditions with a similar magnitude of catch.

STAT response: The requested model uses a stock reduction analysis (SRA) between unfished
conditions and the 1951 start of data. Output is shown in Figure A4. The model had difficulty
converging, and is an approximate solution; also, there are not enough model years to allow
Again, the alternative treatment of initial conditions has an effect mainly on pre-1970 estimates
of abundance, including estimated unfished abundance. Results are not plausible, given the
disagreement with CalCOFI abundance index values the early 1950s. The STAT does not
consider a stock reduction analysis approach for setting initial conditions to be plausible for
bocaccio because it does not recognize that abundance is determined by rare large recruitments.
Because recruitment is sporadic, there is no basis for assuming that the population was at the
theoretical unfished level at any particular year in the past–it could have been anywhere is a
wide range about that value. This bocaccio model is insensitive to these initial conditions, as all
models converge to approximately the same result by 1970. This is partially because the model
tracks only ages 1 to 20, and all recruitments prior to 1950 are collapsed into the “plus-group”
by 1970.

16000
SRA start
14000
Equil C = 1000
12000 Equil C = 2000
spawning output

10000
8000
6000
4000
2000
0
0 20 40 60 80 100 120
year

Figure A4. Trajectory of spawning output resulting from

initializing the model with a stock reduction analysis
from 1930 to 1950, as opposed to assuming equilibrium
under a constant catch.

41
STAR Panel request K (repeats request G). Produce a model with separate blocks of
selectivity for the southern recreational fleet pre and post 2003. Use the working model
(Request C). You may have to extend the last block including earlier years to get convergence.

Reason: Determine the effect of a constant selectivity on the magnitude of recruitment given
that the fishery may have changed due to management.

STAT response: Separate selectivity curves for 2003-2006 were obtained by creating a separate
southern and central California recreational fisheries for that period. Total log likelihood
improved by 13.7 points at a cost of estimating 14 new parameters, which would be rejected by
conventional criteria. The resulting estimate of the 2003 year class strength is larger than
before, contradicting the STAR Panel’s motivation based on a possible overestimate of the 2003
year class due to changes in selectivity. The recreational fishery selectivity curve for central
California indicates greatly decreased selectivity of small fish, presumably because the 2003
year class does not appear in the length compositions.

12000
new
10000
old

8000
Recruitment

6000

4000

2000

0
85

87

89

91

93

95

97

99

101

103

Year

Recl South Recl North

1.2 1.2
1 1

0.8 0.8
Selectivity
Selectivity

old old
0.6 0.6
2004-6 2004-6
0.4 0.4
0.2 0.2
0 0
0 20 40 60 80 0 20 40 60 80
Length Length

42
STAR Panel request L (repeats requests H and G) Determine the co-occurrence of other
species in southern RecFIN trips that caught bocaccio from 1993-2006. For trips that caught
bocaccio, produce the proportion of those trips that also caught other species (only the top 20 or
so).

Reason: Attempt to distinguish near shore from offshore fishing using co-occurrence of species
in the RecFIN source, to confirm no change in recreational fishery selection patterns in recent
years.

STAT response: In southern California, the 2004-2006 pattern of co-occurrences was more like
those of the 1980s and 1990s than was the case for 2000-2003. The STAT disagrees with the
STAR Panel opinion that selectivity changes may account for the large estimated 2003 year
class.

In the following table, a BOLD entry indicates a value higher than in any preceding column,
and a small italic entry indicates a value smaller than in any preceding column.

species\years 1980s 1990s 2000-03 2004 2005 2006

n bocaccio trips (So CA) 200 95 117 385 394 513
Pacific mackerel 0.515 0.295 0.239 0.223 0.305 0.228
kelp bass 0.195 0.095 0.077 0.213 0.180 0.148
barred sandbass 0.125 0.074 0.128 0.130 0.157 0.097
Pacific barracuda 0.085 0.063 0.051 0.073 0.058 0.033
California scorpionfish 0.275 0.242 0.496 0.197 0.246 0.242
ocean whitefish 0.225 0.253 0.333 0.153 0.195 0.175
Pacific bonito 0.180 0.053 0.034 0.101 0.056 0.107
California halibut 0.065 0.011 0.103 0.083 0.071 0.039
California sheephead 0.155 0.168 0.171 0.138 0.140 0.109
vermilion rockfish 0.455 0.453 0.718 0.727 0.652 0.696
white croaker 0.120 0.021 0.085 0.016 0.015 0.035
starry rockfish 0.445 0.295 0.496 0.312 0.343 0.392
treefish 0.070 0.116 0.239 0.091 0.104 0.099
olive rockfish 0.240 0.042 0.145 0.135 0.119 0.136
yellowtail 0.020 0.084 0.017 0.029 0.030 0.023
lingcod 0.175 0.189 0.564 0.452 0.457 0.370
honeycomb rockfish 0.145 0.168 0.274 0.164 0.188 0.183
copper rockfish 0.185 0.284 0.359 0.281 0.234 0.351
flag rockfish 0.425 0.242 0.470 0.270 0.279 0.333
squarespot rockfish 0.345 0.253 0.274 0.127 0.132 0.160
Pacific sanddab 0.190 0.221 0.385 0.195 0.157 0.259
greenspotted rockfish 0.390 0.242 0.410 0.312 0.269 0.316
rosy rockfish 0.265 0.189 0.308 0.151 0.129 0.142

43
Appendix B. Responses to comments and recommendations in 2003 and 2005 STAR
reports.

1.0 Items from Bocaccio 2003 STAR Panel Report

1.1 STAR: Triennial survey selectivity is implausible. The selectivity curve of the triennial
survey appeared nearly uniform over all sizes, which appeared very unlikely for a research
bottom trawl survey.

STAT response: The STAT considers the Triennial Survey to be inappropriate for bocaccio,
because it samples the wrong habitat. The selectivity curve for ages 2+ is flat, but the value for
age 1 (the most pelagic period in the life history) is higher. This is not implausible.

The swept-area abundance estimates can be compared with the 2005 model re-run using only
data north of Pt. Conception, and approximate values of q are shown in the table below. This
provides an equivalent geographic area for comparison; the sum of the two area-specific
models is approximately equal to the single stock model (this comparison is the best we can do
for now). The opinion of the STAT is that the Triennial Survey (and similarly, the NWFSC
combo survey) should be excluded from the assessment model.
Triennial Synthesis est q
year tons CenCal only
1977 13778 18530 0.74
1989 19132 4098 4.67
1992 2584 3225 0.80
1995 1646 2606 0.63
1998 424 2276 0.19
2001 485 3372 0.14
2004 6934 4279 1.62

1.2 STAR: A rebuilding analysis was not brought forward, and was not reviewed. The Panel
provided in this report what it feels are appropriate recommendations for the parameters and
historic recruitments to be used. Specifically, the Panel recommends B1-base and B2 models
with constant recruitment to 1969 as alternative model scenarios (equal probability) with
recruitment resampled only back to 1970. The Panel re-emphasizes its recommendation against
using the Stock Synthesis estimates of steepness or recruitment strength prior to 1970 in
rebuilding analyses.

Biomass and recruitment prior to 1970 are highly uncertain since the only available time series
is the CalCOFI index, which may not be reliable, and in any case would be unable to resolve the
relative strength of individual year-classes.

STAT response: Rebuilding analyses have complied with this recommendation. Values of R/S
were resampled for recruiting years (age 1) since 1970, and have included both B1 and B2
models. Resampling R/S is consistent with the model estimate of steepness ca. 0.2.

44
1.3 STAR: The RecFIN CPUE indices and the triennial survey trends are contradictory.
Fishery- dependent CPUE indices can mask real declines in abundance if fishers are able to
redirect effort to areas of high density. Similarly, the triennial trawl survey may be less efficient
at low stock abundance because bocaccio preferentially occupy untrawlable habitat (varying q
with stock abundance). Generally, the Panel felt that data sources with conflicting information
should not be used together in the assessment.

STAT response: Various STAR Panels have differed on how to treat this kind of conflicting
information. The Council requires a single biomass as the basis of its action. If the assessment
presents two different “equally plausible” numbers, the Council is left with little alternative but to
take a simple average. The STAT considers a model that included both sources of information
(model STATc) to be a better approach than to average the B1 and B2 results. The SSC
concurred with this approach.

1.4 STAR: In general, Stock Synthesis predicted modes within the size composition data for
bocaccio reasonably well, but had a tendency to consistently under-fit the magnitude of the modal
size and overestimate the dispersion about the mode. The residual pattern from the fit to the
length frequency data is unusual and indicates systematic lack of fit. Its effect on the assessment
results is unknown.

STAT response: One promising area of future work would be to develop a time-varying growth
model (similar to that for chilipepper in 2007). This requires migration to the SS2 modeling
framework.

1.5 STAR: Due to the extensive fishery closures and regulations prohibiting retention of catch in
excess of the legal limits, fishery CPUE indices in the future will be biased indices of abundance.
The Council and NMFS need to consider to how to monitor bocaccio status in the future. The
CPFV data set consisting of reef-specific indices of abundance from partyboats is extremely
valuable for evaluating of local fishing effects and as an index of overall abundance. Reef-
specific CPUE is not as subject to the typical limitations of fishery CPUE data. A program of
exempted fishing permits for partyboats with observers to monitor stock status should be
considered.

STAT response: CPUE indexes cease being used in the model after 2003, due to these sorts of
uncertainties. Development of a monitoring program of site-specific CPUE is desirable, but is not
within the scope of the STAT’s capability.

45
1.6 STAR: More attention needs to be given to how growth is modeled in the assessment. A
model with time varying growth or cohort-specific growth may improve the fit to the length
frequency data. Alternative ways to model variation in length with age should also be considered.
Also, the Panel recommends that ageing of bocaccio be re-visited. A modest ageing sample could
be used to evaluate whether the linear trend in the coefficient of variation (CV) of length with age
in Stock Synthesis is a reasonable assumption, as well as confirming the model estimates of
growth.

STAT response: The STAT agrees with the recommendation to develop a variable growth rate
model, which requires use of SS2. Ageing could be revisited if resources were provided to do the
work. Unfortunately, the formal program budget has been insufficient since 2004.

1.7 STAR: The Stock Synthesis model apparently does not perform well with the diverse data
sets used to assess bocaccio. Consideration should be given to moving the bocaccio assessment to
a new modeling environment, ideally one with optimization routines using automatic
differentiation rather than numerical differentiation as in Stock Synthesis.

STAT response: The STAT agrees, though the SS2 model is unlikely in itself to resolve the
problems associated with diverse and in some cases contradictory data sets.

1.8 STAR: Early catch history of bocaccio is a significant source of assessment uncertainty.
Focused research on historical catch is needed. A comprehensive approach should be taken where
historical catches of all West Coast groundfish species are investigated at the same time.
Assessing historical effort in West Coast groundfish fisheries may be more successful as a
collaborative undertaking between an expert in historical research and a stock assessment
scientist.

STAT response: We are making significant progress on this issue, but there is still a long ways to
go The CDFG has now released landings receipt information back to 1969. We have obtained
significant outside funding from the NESDID CDMP program to capture earlier historical landing
data, and have captured summary data by market category (not strictly equivalent to species),
month and geographic block of origin back to 1931. Unfortunately these summaries do not
include gear information. We have microfiche of individual vessel landings data back to 1950,
and work will begin this year to process portions of that massive data set.

1.9 STAR: Work needs to be done to figure how to the start the model with appropriate initial
conditions and with sensible initial depletion which is consistent with the data.

STAT response: The initial conditions in the model are completely plausible, and there is
nothing wrong with them. The assumed background level of annual catch (2000 mtons) is very
close to the estimated MSY values in Table ES4, suggesting that the stock could easily have been
at or below (especially given bocaccio’s tendency toward rare large recruitments) Bmsy at the
beginning of the modeled period, ca. 1950.

46
1.10 STAR: The relationship between the CalCOFI index and climate should be evaluated. Two
analyses are suggested. The first is to compare the residual patterns in model fits to an
environmental index such as the Scripps Pier water temperatures. Adding an environmental
covariate to the CalCOFI index catchability coefficient may improve the model fit to the index if
annual egg production is influenced by environment conditions. A second analysis would be to
compare biomass trends to indices associated with regime-scale environmental variability to see if
significant correlations exist that would help explain long-term abundance trends.

STAT response: Both of these studies are desirable, but are not yet possible due to lack of
information on colder ocean conditions. There was a major warming of California coastal waters
between 1976 and 1998, and there is little sign that temperatures have returned to pre-1976 levels
even yet. Given that the bocaccio model does not produce “reliable” independent estimates
before the early 1970s (when the CalCOFI index itself is the only source of information), we lack
the necessary contrast in ocean conditions to provide the needed information.

Scripps Pier

20
Mean Annual SST

19

18

17

16

15
1915 1935 1955 1975 1995
Year

47
2.0 Items from Bocaccio 2005 STAR Panel Report

2.1 STAR: The triennial survey will likely be discontinued in 2006 so it is desirable to
calibrate the triennial survey indices with those from the NWFSC Combined Survey.

STAT response: The NWFSC has taken the lead on calibration of the trawl surveys, and may
want to consider doing this.

2.2 STAR: Exempted fishing permits are unlikely to provide the quality of catch and effort
data hoped for. If exempted fishing permits are to be used to provide indices of abundance, it is
necessary to check the power of the monitoring program first.

STAT response: This appears to be a response to a recommendation in the 2003 STAR report.

2.3 STAR: An exploratory delta-GLM analysis of the triennial survey was provided to the
STAR Panel. The STAR Panel considered the analysis to be promising and suggested that it be
applied to the NWFSC Combined Survey.

STAT response: The NWFSC is welcome to do this. Generically, they haven’t been willing to
release the data.

2.4 STAR: This species exhibits multiple annual spawning (as a function of age, size, or
environment?). This possibility needs to be investigated based on fish collected from the fisheries
or the survey if an index of spawning output based on larval counts is to be developed for
comparison with the CALCOFI index or juvenile surveys.

STAT response: The STAT concurs. Due to management restrictions, sampling is no longer
encountering enough fish to do this. Funding has been insufficient since 2004.

2.5 STAR: The indices of abundance are assumed to be linearly related to abundance. There
is a possibility of non-linear relationships between the triennial indices and abundance due to
density dependence and habitat (trawlable and untrawlable) considerations. Investigation of
historical data and in situ observations may shed light on some possible relationships.

STAT response: The STAT concurs, though it still may not produce a useful index of abundance.

2.6 STAR: Models with time-varying growth should be included in the assessment if data can
support them. The length data exhibit strong modes which could form the basis for such
estimates.

STAT response: The STAT concurs. This requires SS2.

48
2.7 STAR: Although ageing of bocaccio is difficult, there are large numbers of otoliths that
have been collected, but not been read. There is potential for using the age information to resolve
broad-scale questions regarding changes over time in growth. Multiple reader studies, or other
methods of validation, are desirable to assess reader bias and imprecision.

STAT response: Also see 1.6 above. Funding has been insufficient since 2004.

2.8 STAR: Models could be fitted to data on check marks if there is uncertainty about the
interpretation of check marks as annuli. Check mark data could be treated in the same way as age
data, i.e. subject to ageing bias and ageing imprecision, with the extent of ageing error treated as
estimable within the model.

STAT response: The STAT concurs, but funding has been insufficient since 2004.

2.9 STAR: Future assessments should be based on Stock Synthesis 2. This should allow more
formal quantification of parameter uncertainty. The next assessment should include a formal
comparison of the results of SS1 and SS2 based on the current assessment.

STAT response: The STAT concurs.

2.10 STAR: Consideration should be given to the development of a more spatially disaggregated
model for bocaccio. Although this approach was rejected by the 2002 STAR Panel, improved
CalCOFI coverage north of Pt Conception since 2003 may support more spatial structure within
the assessment.

STAT response: The STAT concurs, with reservations. This requires SS2, which has a limited
capacity to model migration of adults between geographic areas. However, we lack knowledge of
age-specific migration rates, so any model results would be very tentative.

2.11 STAR: According to the STATC model, the spawning output was close to the overfished
threshold in the first year of the model (1951), which differs from the common assumption that
the biomass is close to B0 at the beginning of the analysis. This species has highly variable
recruitment and its biomass would vary substantially over time and a single B0 may not be
appropriate. The STAR Panel stresses the need for guidelines for defining B0 (and hence proxies
for BMSY) for stocks with episodic recruitment. The related problem of what subset of annual
recruitments to average to obtain Recruits/Spawning output values for forecasts should also be
addressed.

STAT response: Also see 1.9 above. The initial conditions are plausible, given the magnitude of
the near-MSY level of assumed background catch. The SS2 model will allow a much earlier start
date, and should be able to portray the dynamics leading to the relatively low 1950 abundance.

49
2.12 STAR: There should be further consideration of the implications of using the prior on
steepness derived by He et al. (in review), including its implications for species with other life
history characteristics.

STAT response: The old Stock Synthesis cannot do this. However, in 2007 the STAT has
included a model (model STATcMPDh) with a stock-recruitment relationship based on the
Maximum Posterior Distribution (MPD) value of steepness (h) from Dorn’s unpublished analysis,
which accomplishes much the same thing.

2.13 STAR: The approach used to estimate B0 for widow rockfish had been modified from the
2003 assessment to be consistent with that on which rebuilding analyses are based (multiplying
average recruitment in the early years of the fishery by unfished spawning biomass per recruit).
This led to a change to the current depletion of 10%. There is a need for more explicit guidance
regarding determination of B0 in assessments and in rebuilding analyses.

STAT response: It is unclear what this means for bocaccio.

2.14 STAR: There is a need for a series of cut-off dates for data to be included in assessments,
with cut-offs dependent on the type of data. The lack of such dates means that assessment authors
may be forced to revise decisions on base-case models very close to the date the assessment needs
to be submitted to the STAR Panel, and even revise the draft assessment after this. Given that
documents are supplied to reviewers two weeks in advance of meetings, major changes in
assessments thereafter could compromise the integrity of the review.

STAT response: Late receipt of critical data was a problem in 2005, but not in 2007.

2.15 STAR: Several of the 2005 assessments have conducted historical catch reconstructions.
An effort needs to be made to develop a consistent approach to reconstructing catch histories. The
ideal outcome would be a single document outlining the best reconstructed catch histories for
each species (c.f. Rogers (2003) that lists foreign catches). The California landing receipts on
microfilm back to 1950 should be incorporated into the landings database.

STAT response: We are currently working on this, with support from the NESDIS/CDMP. It is
not a small job, and may take years.

2.16 STAR: There is still some inconsistency in how assessment authors decide whether to
include or exclude recreational indices in assessments. Attempts to provide guidelines for the
development and use of indices of abundance based on recreational catch and effort data would
be worthwhile.

STAT response: The methodology use in this assessment was presented at a recreational CPUE
workshop, and was endorsed for use.

50
2.17 STAR: Stock Synthesis 2 should be extended to: a) allow assessment authors to include
weight-frequency data in assessments; b) estimate the parameters of the ageing error matrix; and
c) estimate the extent of overdispersion of the indices.

STAT response: When the model migrates to SS2, these features would be useful.

2.18 STAR: The raw data on which recreational length-frequency and catch-effort information
are based should be made available to assessment authors in a convenient format. This will allow
more detailed examination of the spatial patterns, and allow more sophisticated analyses of the
catch-effort information; at present it is impossible to distinguish between lack of data and zero
catch records.

STAT response: Retrieval of recreational data has been an ongoing problem. Some aspects are
“friendly” but others, especially involving disaggregated data can be difficult.

51
Appendix C. Data file for model STATc

2007BocacciodataforCaliforniawithrevisedcommerciallandings
2000,1,trawl,H&L,setnet,recSO,recCEN
50,1,1287,200,0,39,86,1487,125
51,1,1738,277,0,35,98,2015,133
52,1,1691,276,0,45,86,1967,131
53,1,1921,321,0,56,72,2242,128
54,1,1979,337,0,122,91,2316,213
55,1,2034,290,0,213,108,2324,321
56,1,2383,356,0,256,121,2739,377
57,1,2584,365,0,138,120,2949,258
58,1,2621,649,0,95,193,3270,288
59,1,2236,565,0,57,160,2801,217
60,1,2163,351,0,63,125,2514,188
61,1,1631,354,0,72,94,1985,166
62,1,1316,343,0,68,109,1659,177
63,1,1939,386,0,67,111,2325,178
64,1,1229,259,0,94,85,1488,179
65,1,1417,305,0,117,132,1722,249
66,1,2614,332,0,170,142,2946,312
67,1,4325,328,0,210,140,4653,350
68,1,2319,321,0,223,166,2640,389
69,1,1436,304,0,212,154,1740,366
70,1,1660,298,0,289,204,1958,493
71,1,1624,424,0,244,167,2048,411
72,1,2460,598,0,339,226,3058,565
73,1,6033,1040,0,401,260,7073,661
74,1,6968,778,0,459,289,7746,748
75,1,4212,812,0,450,276,5024,726
76,1,3969,776,0,417,248,4745,665
77,1,2172,581,0,377,218,2753,595
78,1,2624,270,125,350,196,3019,546
79,1,3585,371,235,445,242,4191,687
80,1,3641,335,216,1755,178,4192,1933
81,1,3942,314,353,841,230,4609,1071
82,1,4220,395,387,1158,358,5002,1516
83,1,4194,239,588,265,301,5021,566
84,1,3511,369,547,177,67,4427,244
85,1,1231,160,1092,321,66,2483,387
86,1,1165,273,1086,428,171,2524,599
87,1,1206,304,968,90,103,2478,193
88,1,1285,517,371,107,44,2173,151
89,1,1148,386,982,179,78,2516,257
90,1,1144,497,793,233,91,2434,324
91,1,761,271,458,200,92,1490,292
92,1,493,482,640,167,92,1615,259
93,1,560,447,430,109,19,1437,128
94,1,526,207,263,215,5,996,220
95,1,377,69,281,44,3,727,47
96,1,288,93,92,67,26,473,93,,,,
97,1,230,58,35,49,107,323,156,479,,,
98,1,73,42,39,29,23,154,52,206,,,
99,1,45,21,7,71,53,73,124,197,,,
100,1,54,21,2,52,60,77,112,189,,,
101,1,59,35,4,60,49,98,109,207,,,
102,1,41,7,0,76,8,48,84,132,,,
103,1,1,2,0,11,0,3,11,14,,,
104,1,11,9,0.3,60,2,20.3,62,82.3,,,
105,1,23,26,0.1,32,6,49.1,38,87.1,,,
106,1,5,20,0.2,31,11,25.2,42,67.2,,,
-1,1,1,1,1,1,1,1,1,END,OF,CATCH,DATA
-1,1,1,1,END,OF,EFFORT,DATA,,,,,
73,1,7,12,697.38,-826.84,Age1rec'tIndex(powerplants),,,,,,
74,1,7,12,105.92,-107.16,Age1rec'tIndex(powerplants),,,,,,

52
75,1,7,12,228.84,-199.85,Age1rec'tIndex(powerplants),,,,,,
76,1,7,12,266.47,-237.74,Age1rec'tIndex(powerplants)
77,1,7,12,43.85,-43.95,Age1rec'tIndex(powerplants)
78,1,7,12,640.21,-198.21,Age1rec'tIndex(powerplants)
79,1,7,12,116.33,-122,Age1rec'tIndex(powerplants)
80,1,7,12,52.49,-41.53,Age1rec'tIndex(powerplants)
81,1,7,12,31.35,-27.16,Age1rec'tIndex(powerplants)
82,1,7,12,13.48,-14.11,Age1rec'tIndex(powerplants)
83,1,7,12,0.14,-0.29,Age1rec'tIndex(powerplants)
84,1,7,12,0.07,-0.18,Age1rec'tIndex(powerplants)
85,1,7,12,24.75,-22.24,Age1rec'tIndex(powerplants)
86,1,7,12,17.02,-10.4,Age1rec'tIndex(powerplants)
87,1,7,12,6.38,-6.33,Age1rec'tIndex(powerplants)
88,1,7,12,28.11,-41.1,Age1rec'tIndex(powerplants)
89,1,7,12,485.79,-381.52,Age1rec'tIndex(powerplants)
90,1,7,12,9.12,-12.3,Age1rec'tIndex(powerplants)
91,1,7,12,7.56,-5.54,Age1rec'tIndex(powerplants)
92,1,7,12,37.78,-31.77,Age1rec'tIndex(powerplants)
93,1,7,12,5.64,-6.66,Age1rec'tIndex(powerplants)
94,1,7,12,0.5,-1.01,Age1rec'tIndex(powerplants)
95,1,7,12,0.23,-0.47,Age1rec'tIndex(powerplants)
96,1,7,12,8.24,-10.68,Age1rec'tIndex(powerplants)
97,1,7,12,1.69,-2.48,Age1rec'tIndex(powerplants)
98,1,7,12,2.64,-4.51,Age1rec'tIndex(powerplants)
99,1,7,12,0.07,-0.18,Age1rec'tIndex(powerplants)
100,1,7,12,81.81,-111.54,Age1rec'tIndex(powerplants)
101,1,7,12,14.66,-12.72,Age1rec'tIndex(powerplants)
80,1,7,6,0.917,-0.459,MRFnorth
81,1,7,6,1.28,-0.64,MRFnorth
82,1,7,6,1.326,-0.663,MRFnorth
83,1,7,6,1.377,-0.689,MRFnorth
84,1,7,6,0.388,-0.194,MRFnorth
85,1,7,6,0.75,-0.375,MRFnorth
86,1,7,6,1.39,-0.695,MRFnorth
87,1,7,6,0.914,-0.457,MRFnorth
88,1,7,6,0.294,-0.147,MRFnorth
89,1,7,6,0.457,-0.228,MRFnorth
90,1,7,6,-9,-9,Placeholder
91,1,7,6,-9,-9,Placeholder
92,1,7,6,-9,-9,Placeholder
93,1,7,6,0.202,-0.101,MRFnorth
94,1,7,6,0.351,-0.175,MRFnorth
95,1,7,6,0.482,-0.241,MRFnorth
96,1,7,6,0.535,-0.268,MRFnorth
97,1,7,6,0.42,-0.21,MRFnorth
98,1,7,6,0.432,-0.216,MRFnorth
99,1,7,6,0.802,-0.401,MRFnorth
100,1,7,6,1.961,-0.98,MRFnorth
101,1,7,6,2.022,-1.011,MRFnorth
102,1,7,6,2.618,-1.309,MRFnorth
80,1,7,8,3.401,-1.701,MRFsoCAL
81,1,7,8,3.447,-1.724,MRFsoCAL
82,1,7,8,3.173,-1.587,MRFsoCAL
83,1,7,8,1.318,-0.659,MRFsoCAL
84,1,7,8,1.034,-0.517,MRFsoCAL
85,1,7,8,2.224,-1.112,MRFsoCAL
86,1,7,8,1.91,-0.955,MRFsoCAL
87,1,7,8,0.275,-0.137,MRFsoCAL
88,1,7,8,0.169,-0.085,MRFsoCAL
89,1,7,8,0.997,-0.499,MRFsoCAL
90,1,7,8,-9,-9,Placeholder
91,1,7,8,-9,-9,Placeholder
92,1,7,8,-9,-9,Placeholder
93,1,7,8,1.631,-0.81546425,MRFsoCAL
94,1,7,8,1.732,-0.86605425,MRFsoCAL
95,1,7,8,0.448,-0.22416445,MRFsoCAL

53
96,1,7,8,0.246,-0.122946,MRFsoCAL
97,1,7,8,0.395,-0.19748125,MRFsoCAL
98,1,7,8,0.234,-0.11709625,MRFsoCAL
99,1,7,8,0.566,-0.28304295,MRFsoCAL
100,1,7,8,1.098,-0.54898735,MRFsoCAL
101,1,7,8,1.28,-0.63993495,MRFsoCAL
102,1,7,8,2.01,-1.004885,MRFsoCAL
51,1,1,11,0.765,-9,CalCOFIindex
52,1,1,11,1.089,-9,CalCOFIindex
53,1,1,11,0.779,-9,CalCOFIindex
54,1,1,11,1.349,-9,CalCOFIindex
55,1,1,11,1.153,-9,CalCOFIindex
56,1,1,11,0.638,-9,CalCOFIindex
57,1,1,11,1.435,-9,CalCOFIindex
58,1,1,11,1.063,-9,CalCOFIindex
59,1,1,11,0.307,-9,CalCOFIindex
60,1,1,11,0.457,-9,CalCOFIindex
61,1,1,11,0.441,-9,CalCOFIindex
62,1,1,11,0.337,-9,CalCOFIindex
63,1,1,11,0.676,-9,CalCOFIindex
64,1,1,11,0.396,-9,CalCOFIindex
65,1,1,11,0.661,-9,CalCOFIindex
66,1,1,11,1.367,-9,CalCOFIindex
67,1,1,11,-9,-9,Placeholder
68,1,1,11,2.485,-9,CalCOFIindex
69,1,1,11,2.525,-9,CalCOFIindex
70,1,1,11,-9,-9,Placeholder
71,1,1,11,-9,-9,Placeholder
72,1,1,11,1.748,-9,CalCOFIindex
73,1,1,11,-9,-9,Placeholder
74,1,1,11,-9,-9,Placeholder
75,1,1,11,1.106,-9,CalCOFIindex
76,1,1,11,1.317,-9,CalCOFIindex
77,1,1,11,-9,-9,Placeholder
78,1,1,11,0.894,-9,CalCOFIindex
79,1,1,11,-9,-9,Placeholder
80,1,1,11,-9,-9,Placeholder
81,1,1,11,0.703,-9,CalCOFIindex
82,1,1,11,-9,-9,Placeholder
83,1,1,11,-9,-9,Placeholder
84,1,1,11,0.497,-9,CalCOFIindex
85,1,1,11,0.089,-9,CalCOFIindex
86,1,1,11,0.134,-9,CalCOFIindex
87,1,1,11,0.683,-9,CalCOFIindex
88,1,1,11,0.597,-9,CalCOFIindex
89,1,1,11,0.472,-9,CalCOFIindex
90,1,1,11,0.336,-9,CalCOFIindex
91,1,1,11,0.467,-9,CalCOFIindex
92,1,1,11,0.592,-9,CalCOFIindex
93,1,1,11,0.107,-9,CalCOFIindex
94,1,1,11,0.169,-9,CalCOFIindex
95,1,1,11,0.062,-9,CalCOFIindex
96,1,1,11,1.088,-9,CalCOFIindex
97,1,1,11,0.218,-9,CalCOFIindex
98,1,1,11,0.082,-9,CalCOFIindex
99,1,1,11,0.129,-9,CalCOFIindex
100,1,1,11,0.121,-9,CalCOFIindex
101,1,1,11,0.065,-9,CalCOFIindex
102,1,1,11,0.301,-9,CalCOFIindex
103,1,1,11,0.403,-9,CalCOFIindex
104,1,1,11,0.376,-9,CalCOFIindex
105,1,1,11,0.358,-9,CalCOFIindex
106,1,1,11,0.53,-9,CalCOFIindex
77,1,7,10,999,-9,1977,TRIENNIAL,
78,1,7,10,-9,-9,Placeholder,,
79,1,7,10,-9,-9,Placeholder,,

54
80,1,7,10,691,-9,1980,TRIENNIAL,
81,1,7,10,-9,-9,Placeholder,,
82,1,7,10,-9,-9,Placeholder,,
83,1,7,10,1181.5,-9,1983,TRIENNIAL,INDEX
84,1,7,10,-9,-9,Placeholder,,
85,1,7,10,-9,-9,Placeholder,,
86,1,7,10,637.5,-9,1986,TRIENNIAL,INDEX
87,1,7,10,-9,-9,Placeholder,,
88,1,7,10,-9,-9,Placeholder,,
89,1,7,10,735.5,-9,1989,TRIENNIAL,INDEX
90,1,7,10,-9,-9,Placeholder,,
91,1,7,10,-9,-9,Placeholder,,
92,1,7,10,186,-9,1992,TRIENNIAL,INDEX
93,1,7,10,-9,-9,Placeholder,,
94,1,7,10,-9,-9,Placeholder,,
95,1,7,10,82.7,-9,1995,TRIENNIAL,INDEX
96,1,7,10,-9,-9,Placeholder,,
97,1,7,10,-9,-9,Placeholder,,
98,1,7,10,16.7,-9,1998,TRIENNIAL,INDEX
99,1,7,10,-9,-9,Placeholder,,
100,1,7,10,-9,-9,Placeholder,,
101,1,7,10,34,-9,2001,TRIENNIAL,INDEX
102,1,7,10,-9,-9,Placeholder,,
103,1,7,10,-9,-9,Placeholder,,
104,1,7,10,274.6,-9,2004,TRIENNIAL,INDEX
84,1,7,13,0.004,-0.002,JuvSurveyrectmt,,
85,1,7,13,17.384,-8.692,JuvSurveyrectmt,,
86,1,7,13,0.004,-0.002,JuvSurveyrectmt,,
87,1,7,13,0.695,-0.3475,JuvSurveyrectmt,,
88,1,7,13,0.994,-0.497,JuvSurveyrectmt
89,1,7,13,1.095,-0.5475,JuvSurveyrectmt
90,1,7,13,0.182,-0.091,JuvSurveyrectmt
91,1,7,13,0.091,-0.0455,JuvSurveyrectmt
92,1,7,13,0.515,-0.2575,JuvSurveyrectmt
93,1,7,13,0.002,-0.001,JuvSurveyrectmt
94,1,7,13,0.129,-0.0645,JuvSurveyrectmt
95,1,7,13,0.007,-0.0035,JuvSurveyrectmt
96,1,7,13,0.013,-0.0065,JuvSurveyrectmt
97,1,7,13,0.004,-0.002,JuvSurveyrectmt
98,1,7,13,0.018,-0.009,JuvSurveyrectmt
99,1,7,13,0.004,-0.002,JuvSurveyrectmt
100,1,7,13,0.027,-0.0135,JuvSurveyrectmt
101,1,7,13,0.051,-0.0255,JuvSurveyrectmt
102,1,7,13,0.079,-0.0395,JuvSurveyrectmt
103,1,7,13,0.342,-0.171,JuvSurveyrectmt
82,1,7,9,166.4,-83.2,areaweightedCPUEfromRalston
83,1,7,9,73.1,-36.55,areaweightedCPUEfromRalston
84,1,7,9,72.3,-36.15,areaweightedCPUEfromRalston
85,1,7,9,30.7,-15.35,areaweightedCPUEfromRalston
86,1,7,9,31.2,-15.6,areaweightedCPUEfromRalston
87,1,7,9,44.4,-22.2,areaweightedCPUEfromRalston
88,1,7,9,51.6,-25.8,areaweightedCPUEfromRalston
89,1,7,9,35.8,-17.9,areaweightedCPUEfromRalston
90,1,7,9,37.1,-18.55,areaweightedCPUEfromRalston
91,1,7,9,26.9,-13.45,areaweightedCPUEfromRalston
92,1,7,9,20.4,-10.2,areaweightedCPUEfromRalston
93,1,7,9,19.7,-9.85,areaweightedCPUEfromRalston
94,1,7,9,23.9,-11.95,areaweightedCPUEfromRalston
95,1,7,9,15.2,-7.6,areaweightedCPUEfromRalston
96,1,7,9,8.7,-4.35,areaweightedCPUEfromRalston
87,1,7,7,3.545,-1.7725,VandenbergCPUE
88,1,7,7,2.349,-1.1745,VandenbergCPUE
89,1,7,7,3.001,-1.5005,VandenbergCPUE
90,1,7,7,6.009,-3.0045,VandenbergCPUE
91,1,7,7,4.637,-2.3185,VandenbergCPUE
92,1,7,7,3.543,-1.7715,VandenbergCPUE

55
93,1,7,7,2.319,-1.1595,VandenbergCPUE
94,1,7,7,1.46,-0.73,VandenbergCPUE
95,1,7,7,1.721,-0.8605,VandenbergCPUE
96,1,7,7,1.457,-0.7285,VandenbergCPUE
97,1,7,7,1.823,-0.9115,VandenbergCPUE
98,1,7,7,1.646,-0.823,VandenbergCPUE
81,1,7,14,33.058,-16.529,MRFpierRectmt
82,1,7,14,2.807,-1.4035,MRFpierRectmt
83,1,7,14,0.003,-0.0015,MRFpierRectmt
84,1,7,14,0.005,-0.0025,MRFpierRectmt
85,1,7,14,43.127,-21.5635,MRFpierRectmt
86,1,7,14,6.987,-3.4935,MRFpierRectmt
87,1,7,14,0.498,-0.249,MRFpierRectmt
88,1,7,14,13.529,-6.7645,MRFpierRectmt
89,1,7,14,77.056,-38.528,MRFpierRectmt
90,1,7,14,1.081,-0.5405,MRFpierRectmt
91,1,7,14,-9,-9,Placeholder
92,1,7,14,-9,-9,Placeholder
93,1,7,14,-9,-9,Placeholder
94,1,7,14,18.623,-9.3115,MRFpierRectmt
95,1,7,14,0.003,-0.0015,MRFpierRectmt
96,1,7,14,0.312,-0.156,MRFpierRectmt
97,1,7,14,0.13,-0.065,MRFpierRectmt
98,1,7,14,0.003,-0.0015,MRFpierRectmt
99,1,7,14,0.003,-0.0015,MRFpierRectmt
100,1,7,14,0.105,-0.0525,MRFpierRectmt
101,1,7,14,0.003,-0.0015,MRFpierRectmt
102,1,7,14,0.003,-0.0015,MRFpierRectmt,,,,,,,,,,,,,,,,,,,
103,1,7,14,0.003,-0.0015,MRFpierRectmt,,,,,,,,,,,,,,,,,,,
-1,1,1,1,1,1,END,OF,,,,,,,,,,,,,,,,,,
-1,-1,<==,No,aging,error(not,used),,,,,,,,,,,,,,,,,,,
-1,-1,,,,,,,,,,,,,,,,,,,,,,,,
-1,-1,,,,,,,,,,,,,,,,,,,,,,,,
25,25,"<==25lengthbins24..68at2cm,72,76 bins",,,,,,,,,,,,,,,,,,,,,,,
24,26,28,30,32,34,36,38,40,42,44,46,48,50,52,54,56,58,60,62,64,66,68,72,76,80
47.6,-0.2876,length@50%matureslopeEcheverria1987,,,,,,,,,,,,,,,,,,,,,,,
6.17E-06,3.1712,Length-weightparsfemale1995TriennialTrawl(Ralston),,,,,,,,,,,,,,,,,,,,,,,
0.22475,0.03657,eggs/kginterceptandslopeReinterpretedfromPhillipsbyRalston1996,,,,,,,,,,,,,,,,,,,,,,,
6.17E-06,3.1712,Length-weightparsmale1995TriennialTrawl(Ralston),,,,,,,,,,,,,,,,,,,,,,,
YEAR,PER,TYPE,KIND,MAXSEX,TOTAGED,MIN1,MIN2,MAX1,MAX2,MARKET,,,,,,,,,,,,,,,
75,1,4,4,0,157,1,1,25,25,0,nfish=,21486,,,,,,,,,,,,,
136,1199,2795,1908,1664,3328,3599,2204,826,502,584,765,691,455,311,203,110,71,52,36,17,9,13,7,1,
76,1,4,4,0,173,1,1,25,25,0,nfish=,26209,,,,,,,,,,,,,
151,457,781,545,625,2751,4173,2594,3197,3597,2066,1087,985,1003,820,518,297,212,129,93,52,29,32,14,1
77,1,4,4,0,122,1,1,25,25,0,nfish=,11155,,,,,,,,,,,,
54,88,138,93,208,424,484,432,1011,1645,1570,1535,1047,611,566,428,332,177,106,72,60,42,24,7,1
77,1,10,4,3,0,1,1,25,25,0,nfish=,,nsamps=,30,,,,,,,,,,
2100,0,1088,1088,8225,26005,35918,154731,161624,170535,138161,93622,111977,44689,48380,104669,60728,98818,66653,112582,70692,665
36,119451,11354,637
6583,2702,4354,4779,14761,20887,44556,79087,227801,190667,131989,102300,79657,92392,100508,174131,106070,189490,106751,134337,4
4918,11575,0,0,0
78,1,1,4,3,106,1,1,25,25,0,nfish=,1565,nsamps=,142,,,,,,,,,,
100,121,585,4005,6572,4236,2302,1640,9773,3363,13568,13662,42582,41869,36318,18511,14589,9568,23918,21089,13940,7623,14640,13339,
7477
0,0,74,1675,892,2802,3004,6250,5968,13768,39199,62849,51166,30362,25922,10772,22040,19771,14616,10438,3286,3355,972,603,603
78,1,3,4,3,19,1,1,25,25,0,nfish=,61,nsamps=,6,,,,,,,,,,
0,0,0,0,0,0,0,0,417,476,441,900,494,763,999,685,209,232,232,166,232,122,607,209,163
0,0,0,0,0,0,0,0,0,166,209,288,1508,1021,859,807,209,209,456,0,0,122,0,0,122
78,1,4,4,0,145,1,1,25,25,0,nfish=,17988,,,,,,,,,,,,
2046,3184,2073,552,125,199,299,272,500,870,1084,1360,1414,1220,914,655,457,325,210,114,45,35,27,6,2
79,1,1,4,3,104,1,1,25,25,0,nfish=,1448,nsamps=,102,,,,,,,,,,
0,0,0,0,1108,2883,28218,105365,22315,2141,13913,13913,389,17719,105814,61823,19433,1996,22315,46172,614,2630,6620,1821,1013
0,0,0,0,700,15142,25270,25032,0,23061,0,758,70685,118299,44871,19611,42608,84105,14990,17943,8853,1292,700,2186,132
80,1,1,4,3,108,1,1,25,25,0,nfish=,1673,nsamps=,225,,,,,,,,,,
0,0,0,10142,11618,10534,10473,62228,244551,308435,228392,70611,19166,19756,60228,66162,42242,29128,22454,31675,27028,18012,42322,
7925,361

56
0,5071,0,0,12622,24720,31673,108613,266944,232919,70825,48886,81575,57566,65004,33864,67178,9899,20704,16301,1543,0,752,0,0
80,1,2,4,3,3,1,1,25,25,0,nfish=,30,nsamps=,2,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,0,0,0,0,0,0,1607,0,0,4821,388,4821,2383,5209,1607
0,0,0,0,0,0,0,0,0,0,0,0,0,0,1607,0,1607,6428,8035,388,1995,0,0,0,0
80,1,4,4,0,92,1,1,25,25,0,nfish=,2577,,,,,,,,,,,,
55,67,75,63,73,105,232,517,524,258,113,77,72,83,80,61,48,39,18,7,4,5,1,0,0
80,1,5,4,0,45,1,1,25,25,0,nfish=,250,,,,,,,,,,,,
5,10,3,6,0,1,9,22,25,17,18,12,15,18,13,11,9,7,12,6,6,10,6,5,4
80,1,10,4,3,0,1,1,25,25,0,nfish=,,nsamps=,17,,,,,,,,,,
33117,93977,33116,0,0,0,25548,223786,540038,730159,489799,141297,0,65385,24126,36625,0,32693,1966,22160,0,0,0,0,0
33116,146555,57954,8279,0,0,53971,254433,827132,761859,270912,32441,1966,11185,24126,3567,65386,98731,54853,28300,21256,0,0,0,0
81,1,1,4,3,101,1,1,25,25,0,nfish=,1290,nsamps=,160,,,,,,,,,,
0,0,0,8132,15419,10123,0,9428,38669,66076,110869,224391,271337,137066,7854,1291,9356,20144,11479,6821,5277,27488,13201,16702,780
0,0,0,4148,1551,4207,47800,68793,90004,161622,173418,126448,68308,63466,33931,25411,43006,27675,51709,7999,7098,3184,10855,0,0
81,1,4,4,0,91,1,1,25,25,0,nfish=,2227,,,,,,,,,,,,
7,22,26,61,146,261,267,179,158,157,215,265,122,78,67,67,48,40,21,9,6,1,1,3,0
81,1,5,4,0,45,1,1,25,25,0,nfish=,250,,,,,,,,,,,,
0,0,1,1,3,2,13,10,6,27,40,30,22,6,6,13,13,13,9,6,8,6,8,5,2
82,1,1,4,3,122,1,1,25,25,0,nfish=,2399,nsamps=,242,,,,,,,,,,
0,0,21,21,245,111,2107,32901,31959,46688,63213,40021,60016,169057,145053,209144,19139,6476,14085,19319,17509,12616,24086,54532,13
18
0,0,7,7,682,155,8412,35822,120468,58028,52391,139363,165215,67210,33173,20159,27226,56484,33410,24561,7645,54,0,0,0
82,1,2,4,3,3,1,1,25,25,0,nfish=,19,nsamps=,2,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,0,0,7237,21711,0,7237,13987,6750,13500,6750,13987,0,6750,6750,0
0,0,0,0,0,0,0,0,0,0,6750,0,0,0,0,0,0,0,13500,0,0,0,6750,0,0
82,1,4,4,0,90,1,1,25,25,0,nfish=,1828,,,,,,,,,,,,
1,2,9,18,36,39,61,156,211,218,214,187,224,176,112,70,45,22,11,5,7,0,3,1,0
82,1,5,4,0,55,1,1,25,25,0,nfish=,310,,,,,,,,,,,,
0,0,0,0,3,5,4,9,15,12,10,25,47,43,49,29,19,13,5,9,7,2,4,0,0
83,1,1,4,3,128,1,1,25,25,0,nfish=,2675,nsamps=,308,,,,,,,,,,
0,0,0,0,101,0,879,939,2635,29438,44537,58133,52133,51175,82114,111799,129765,37199,24640,11723,19779,21341,32899,57707,10927
0,0,449,71,0,258,623,2075,4027,15157,39981,70037,86302,90871,74135,39829,34316,16150,29115,8781,13600,100,202,0,0
83,1,2,4,3,7,1,1,25,25,0,nfish=,55,nsamps=,5,,,,,,,,,,
0,0,0,0,0,0,0,0,304,304,0,608,912,1207,2702,2560,1414,3382,0,903,259,259,903,0,0
0,0,0,0,0,0,0,0,304,0,2702,563,304,1718,2488,1326,1725,2560,0,1790,0,0,0,0,0
83,1,3,4,3,18,1,1,25,25,0,nfish=,44,nsamps=,7,,,,,,,,,,
0,0,0,0,0,0,0,0,2364,4774,7746,3120,12516,5382,2912,10404,9856,0,0,0,4728,0,2410,0,0
0,0,0,0,0,0,0,0,3120,0,6908,12799,3718,4774,13378,13352,6763,1456,0,0,0,0,0,0,0
83,1,4,4,0,86,1,1,25,25,0,nfish=,706,,,,,,,,,,,,
0,2,0,2,9,20,51,73,63,61,83,56,51,43,50,46,33,21,12,13,8,2,6,1,0
83,1,5,4,0,64,1,1,25,25,0,nfish=,359,,,,,,,,,,,,
0,0,2,3,4,4,1,4,6,8,19,27,40,45,52,47,37,22,14,9,8,4,1,0,2
83,1,10,4,3,0,1,1,25,25,0,nfish=,,nsamps=,15,,,,,,,,,,
0,0,0,0,0,5559,0,2590,7260,18905,25146,40713,88899,60051,200335,377143,447870,99634,6881,13005,9991,0,23761,13346,0
0,0,0,0,0,0,11118,0,0,41461,48905,50500,146252,499926,457074,114536,105259,102367,31229,19404,6947,0,0,0,0
84,1,1,4,3,126,1,1,25,25,0,nfish=,2603,nsamps=,276,,,,,,,,,,
0,0,0,0,0,27,4,1222,997,4350,4593,5385,6391,19206,17669,21232,30809,34952,19181,7068,7117,4547,8704,13830,2149
0,0,0,0,0,334,130,1155,3075,8964,7765,12360,28371,63068,42630,32984,13427,9467,19091,6474,10536,1279,1143,0,0
84,1,2,4,3,3,1,1,25,25,0,nfish=,34,nsamps=,2,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,0,0,0,0,0,2659,0,5909,0,2659,4875,3250,1034,2659,0
0,0,0,0,0,0,0,0,0,0,0,0,2068,1034,7977,3102,3102,1034,2659,0,0,0,0,0,0
84,1,3,4,3,18,1,1,25,25,0,nfish=,44,nsamps=,7,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,0,0,0,679,248,1003,517,196,321,642,321,0,0,147,0
0,0,0,0,0,0,0,0,0,0,196,0,1318,1251,789,856,1713,196,0,388,0,0,0,0,0
84,1,4,4,0,85,1,1,25,25,0,nfish=,481,,,,,,,,,,,,
2,3,4,3,8,8,6,21,31,40,57,70,63,48,44,28,16,13,5,3,6,0,2,0,0
84,1,5,4,0,33,1,1,25,25,0,nfish=,183,,,,,,,,,,,,
0,0,0,1,2,2,9,9,9,12,15,17,15,14,23,11,8,7,11,4,6,5,1,1,1
85,1,1,4,3,108,1,1,25,25,0,nfish=,1658,nsamps=,262,,,,,,,,,,
0,20,106,1224,1354,826,312,55,259,805,558,1425,1422,5282,4758,4550,5330,13190,14374,4732,2161,1023,2133,1445,185
0,41,62,338,1291,864,371,729,228,851,2588,4546,4477,11444,21066,17782,8281,2438,2559,1054,847,108,0,0,0
85,1,2,4,3,5,1,1,25,25,0,nfish=,34,nsamps=,4,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,0,54,54,390,216,789,399,27,27,0,0,237,135,0,0
0,0,0,0,0,0,0,0,0,0,0,0,249,214,1531,264,1024,0,0,0,0,0,0,0,0
85,1,3,4,3,29,1,1,25,25,0,nfish=,274,nsamps=,38,,,,,,,,,,
0,0,0,37,0,0,0,0,0,810,159,508,167,2220,2268,2095,2694,4699,15713,11258,4965,1144,6781,5688,0

57
0,0,0,37,0,0,0,0,0,0,1486,5206,3244,3108,14332,11954,22873,3434,9025,3900,1242,2648,0,0,0
85,1,4,4,0,88,1,1,25,25,0,nfish=,1256,,,,,,,,,,,,
126,244,246,151,81,22,8,15,11,22,32,43,57,57,41,43,29,13,14,1,0,0,0,0,0
85,1,5,4,0,95,1,1,25,25,0,nfish=,532,,,,,,,,,,,,
45,103,98,44,7,11,7,6,8,13,15,17,20,16,24,12,14,22,12,16,9,6,4,3,0
86,1,1,4,3,123,1,1,25,25,0,nfish=,2431,nsamps=,189,,,,,,,,,,
0,0,0,2088,11803,21288,23311,23906,12987,4475,2028,121,862,1237,1970,2127,4140,6105,4698,5111,3827,1312,820,861,0
0,145,90,1205,11842,26146,31768,19894,9987,3879,422,1688,984,3245,10560,14182,12740,7080,1763,1830,305,118,40,0,0
86,1,2,4,3,42,1,1,25,25,0,nfish=,496,nsamps=,32,,,,,,,,,,
0,0,156,0,0,491,10,509,647,608,398,10,305,1206,395,1457,1106,1164,1663,493,461,0,0,0,0
0,0,0,33,46,132,756,242,242,23,198,100,1561,1259,1444,1870,668,9,409,0,0,0,0,0,0
86,1,3,4,3,91,1,1,25,25,0,nfish=,1566,nsamps=,152,,,,,,,,,,
0,0,0,0,1800,10,10,250,976,2775,162,58,3662,5472,5398,11705,7989,9269,6816,3860,2395,1025,2480,1601,21
0,0,0,0,0,0,265,2553,272,278,2697,325,6363,14720,15423,10100,10174,5107,5199,1413,1896,0,910,0,0
86,1,4,4,0,88,1,1,25,25,0,nfish=,1267,,,,,,,,,,,,
27,54,45,38,96,223,350,208,54,10,10,20,23,23,20,23,21,4,8,5,3,1,0,1,0
86,1,5,4,0,168,1,1,25,25,0,nfish=,942,,,,,,,,,,,,
8,19,13,25,64,167,245,167,47,4,2,8,13,14,15,16,19,27,16,17,12,12,8,3,1
86,1,10,4,3,0,1,1,25,25,0,nfish=,,nsamps=,17,,,,,,,,,,
11876,5938,2969,2969,39191,43313,45486,48763,19703,0,0,0,0,0,30877,30877,0,17774,70640,154383,255899,154383,185260,0,30877
2969,50738,32197,0,55012,68986,140922,58386,18851,0,0,0,0,0,70640,101516,51786,70640,61754,61754,48650,0,0,0,0
87,1,1,4,3,132,1,1,25,25,0,nfish=,2876,nsamps=,200,,,,,,,,,,
0,0,0,0,124,4311,7620,11188,25074,46838,50776,26040,4086,3077,635,1279,987,1327,2627,1811,4826,3336,1284,633,147
0,0,0,421,1988,6062,9338,24878,45343,49617,29868,10514,1573,2702,2457,4427,9931,4691,3114,1325,196,483,0,0,0
87,1,2,4,3,29,1,1,25,25,0,nfish=,274,nsamps=,22,,,,,,,,,,
0,0,0,85,0,2457,289,275,354,2126,3746,1154,252,768,1255,1922,1445,755,283,17,691,0,0,0,0
0,0,0,0,36,81,7605,15561,8827,6034,5064,162,4566,8823,951,231,2551,220,4914,2457,2457,2457,2457,0,0
87,1,3,4,3,73,1,1,25,25,0,nfish=,1193,nsamps=,101,,,,,,,,,,
0,0,0,0,0,0,0,366,540,4725,11168,5989,2851,1627,3033,7337,6295,6545,2814,1545,1690,689,1738,72,553
0,0,0,0,0,0,984,161,9853,9354,6880,6260,18255,5464,17485,11960,9145,8437,679,1099,944,12,0,0,0
87,1,4,4,0,84,1,1,25,25,0,nfish=,121,,,,,,,,,,,,
3,8,12,2,5,8,14,14,16,8,6,8,2,4,3,4,3,1,0,0,0,0,0,0,0
87,1,5,4,0,203,1,1,25,25,0,nfish=,1136,,,,,,,,,,,,
6,3,6,11,26,69,138,132,110,118,124,75,41,29,20,42,29,29,24,42,22,7,24,7,2
88,1,1,4,3,111,1,1,25,25,0,nfish=,1822,nsamps=,165,,,,,,,,,,
0,11,11,533,1811,1882,2058,8531,11862,15817,21983,27929,31081,28542,11959,4051,599,1221,1608,5593,3840,1634,1037,213,163
0,0,0,83,176,2898,6332,17566,24250,26521,43257,62601,25899,9422,4540,7384,4288,5277,2190,677,268,21,1,0,0
88,1,2,4,3,13,1,1,25,25,0,nfish=,147,nsamps=,10,,,,,,,,,,
0,0,0,0,0,0,0,5184,90,90,18396,18959,32802,17482,9106,0,619,146,0,0,0,0,0,0,0
0,0,0,0,0,0,0,64,0,5771,25977,12757,619,176,1070,32,0,507,0,0,0,0,0,0,0
88,1,3,4,3,73,1,1,25,25,0,nfish=,1189,nsamps=,86,,,,,,,,,,
0,0,0,0,0,254,0,0,159,630,1988,3833,5465,2144,1381,858,1792,944,872,619,404,145,492,0,0
0,0,0,0,0,14,14,47,340,2048,2721,3800,2623,1594,1548,2052,718,238,113,75,2,0,14,0,0
88,1,4,4,0,79,1,1,25,25,0,nfish=,79,,,,,,,,,,,,
0,7,3,3,1,2,8,10,11,6,8,6,4,5,3,1,0,1,0,0,0,0,0,0,0
88,1,5,4,0,226,1,1,25,25,0,nfish=,1264,,,,,,,,,,,,
6,15,11,11,16,31,82,58,107,191,153,157,129,82,41,18,16,27,24,20,15,20,26,6,2
89,1,1,4,3,98,1,1,25,25,0,nfish=,1112,nsamps=,141,,,,,,,,,,
0,0,331,329,10709,13605,27916,29761,10987,3934,5524,23868,8767,16543,22295,14959,6107,10646,787,1018,283,3190,3059,100,261
0,87,2587,12946,10575,7350,21766,13700,37105,18034,12488,35679,36298,17885,12248,1075,9663,6885,5476,1179,531,618,0,0,0
89,1,2,4,3,31,1,1,25,25,0,nfish=,399,nsamps=,24,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,265,264,612,2945,228,7830,5265,326,122,2584,2538,0,0,0,0
0,0,0,0,0,0,0,0,0,0,655,1091,66,5500,2750,76,579,76,0,33,0,0,46,0,0
89,1,3,4,3,87,1,1,25,25,0,nfish=,1486,nsamps=,128,,,,,,,,,,
0,0,0,8,0,704,23,21,632,721,3434,4675,9141,12679,5826,4483,2209,1384,1796,120,189,28,327,17,0
0,0,0,4,6,0,0,0,641,3153,5035,15446,17752,6458,7172,2397,2102,652,243,200,29,1,19,0,0
89,1,4,4,0,85,1,1,25,25,0,nfish=,478,,,,,,,,,,,,
17,60,59,57,24,9,16,20,27,22,29,26,29,19,19,8,6,9,6,5,1,6,1,2,1
89,1,5,4,0,274,1,1,25,25,0,nfish=,1537,,,,,,,,,,,,
37,68,68,20,36,83,91,45,40,65,126,159,177,162,125,74,57,18,25,14,13,9,14,8,3
89,1,10,4,3,47,1,1,25,25,0,nfish=,,nsamps=,69,,,,,,,,,,
876680,6202302,8717749,1081340,287569,652240,534494,113229,2695,6886,2054,0,17223,47746,17018,50518,16642,10211,7604,0,8370,4498
,2230,0,0
927303,7139287,7574159,647602,277457,312915,330241,26151,0,88036,5344,48693,59395,80266,29019,6725,12441,22652,12441,0,0,0,0,0,0
90,1,1,4,3,117,1,1,25,25,0,nfish=,2133,nsamps=,188,,,,,,,,,,
0,0,5,98,1315,7277,11952,17263,17129,7194,3842,4812,2279,5290,4302,6080,11025,4644,2953,1463,655,953,4591,1089,79

58
0,0,86,543,3429,12424,23668,31738,8144,10790,5094,2894,8864,16498,17603,11875,7040,6695,3454,1562,1685,477,0,0,0
90,1,2,4,3,13,1,1,25,25,0,nfish=,141,nsamps=,10,,,,,,,,,,
0,0,0,0,0,0,0,2656,5004,0,1422,1649,2552,2305,774,5552,723,977,0,0,0,926,0,0,0
0,0,0,0,0,0,2254,4508,6332,3071,3501,2051,3845,3795,1649,977,51,0,0,0,0,0,0,0,0
90,1,3,4,3,61,1,1,25,25,0,nfish=,950,nsamps=,105,,,,,,,,,,
0,0,0,0,3978,1,34,372,761,179,301,862,17457,19883,15798,25954,5225,355,1952,5598,4892,67,629,0,0
0,6,0,0,0,182,278,429,312,125,1818,17341,15870,13072,1405,2322,1744,3981,2,0,0,0,0,0,0
90,1,5,4,0,174,1,1,25,25,0,nfish=,974,,,,,,,,,,,,
1,6,11,26,80,161,143,57,61,78,50,45,41,56,52,42,29,11,12,4,2,1,5,0,0
91,1,1,4,3,125,1,1,25,25,0,nfish=,2525,nsamps=,117,,,,,,,,,,
0,0,114,331,424,2091,4515,16776,42312,48563,34893,17372,15795,6347,3982,5700,9435,7219,5124,1843,2243,381,928,298,225
0,66,310,243,361,669,9095,32577,61578,55524,34750,18199,9964,11308,9230,8395,7021,4214,6394,2691,1120,81,265,0,0
91,1,2,4,3,35,1,1,25,25,0,nfish=,253,nsamps=,27,,,,,,,,,,
0,0,0,0,0,323,0,0,194,1383,2000,699,862,506,283,781,607,1056,2758,585,288,0,400,288,0
0,0,50,0,0,0,0,704,1205,3372,938,1617,659,800,842,1662,740,272,628,0,448,288,0,0,0
91,1,3,4,3,40,1,1,25,25,0,nfish=,508,nsamps=,36,,,,,,,,,,
0,0,0,0,0,16,16,123,2438,13494,10260,6334,3365,1251,2899,77,3593,1151,535,16,0,16,823,0,0
0,0,0,0,0,16,46,382,2716,3775,2252,1866,1123,5038,3263,273,294,794,0,6,0,0,0,0,0
91,1,5,4,0,155,1,1,25,25,0,nfish=,866,,,,,,,,,,,,
5,5,4,3,4,13,17,38,93,123,84,63,55,56,58,62,43,38,34,24,14,9,14,6,1
92,1,1,4,3,108,1,1,25,25,0,nfish=,1630,nsamps=,70,,,,,,,,,,
0,0,145,838,2856,5080,4441,4267,2462,3811,11781,23290,20918,13366,7213,5697,5406,4004,7611,3585,4932,3150,3074,2086,38
0,0,0,992,2839,825,5643,3938,4763,18121,23572,22580,14385,9024,7138,7697,4616,4733,6056,3007,367,484,60,0,0
92,1,2,4,3,51,1,1,25,25,0,nfish=,559,nsamps=,39,,,,,,,,,,
0,0,0,0,0,1136,1303,3541,77,2440,3283,12842,11621,5534,1288,1322,554,389,647,285,925,0,775,0,314
0,0,0,0,0,495,1898,501,1274,5320,12452,9265,2475,2711,1456,1444,455,813,620,612,235,153,0,0,0
92,1,3,4,3,76,1,1,25,25,0,nfish=,1258,nsamps=,59,,,,,,,,,,
0,0,0,0,0,36,173,293,713,818,9066,19287,15963,9453,5288,3271,2259,1835,1439,1641,0,19,0,0,0
0,0,0,0,0,48,68,1253,1017,6541,13885,10935,5124,4231,6115,2324,1052,95,25,0,0,0,0,0,0
92,1,5,4,0,303,1,1,25,25,0,nfish=,1697,,,,,,,,,,,,
15,9,12,10,46,85,42,36,71,101,153,230,204,142,106,87,57,84,83,49,26,15,19,11,4
92,1,10,4,3,24,1,1,25,25,0,nfish=,,nsamps=,35,,,,,,,,,,
7363,35287,59359,42417,35617,72287,23500,26438,25188,69265,147767,143856,136155,24273,0,12594,6297,10591,0,0,4117,14428,0,0,0
15414,66542,68362,17115,33038,59154,111627,38610,97391,119640,144828,89281,32578,51149,6297,11742,14428,0,0,0,2043,0,0,0,0
93,1,1,4,3,107,1,1,25,25,0,nfish=,1615,nsamps=,68,,,,,,,,,,
0,0,635,8891,11973,14930,3727,13386,14419,7670,10369,9260,13273,10889,15173,7910,7069,4139,3404,1742,1224,1448,3855,0,0
0,0,50,9015,16719,9836,9560,10853,13509,12739,17169,19556,17727,8444,11149,9586,3436,2809,1651,904,123,0,0,0,0
93,1,2,4,3,80,1,1,25,25,0,nfish=,712,nsamps=,61,,,,,,,,,,
0,0,0,0,0,175,0,55,233,314,218,3381,4985,2131,1962,366,566,716,328,1076,75,302,1041,307,60
0,0,0,116,55,116,110,203,368,1786,3105,836,1570,920,1256,244,1523,1910,488,240,0,0,0,0,0
93,1,3,4,3,60,1,1,25,25,0,nfish=,924,nsamps=,44,,,,,,,,,,
0,0,0,62,50,10,223,400,658,1428,6907,14718,9157,3401,3944,3434,2601,3521,3097,1186,112,568,0,0,0
0,0,0,0,34,20,133,631,1000,5553,5909,6107,3835,3040,2416,2431,228,0,0,0,0,0,0,0,0
93,1,4,4,0,84,1,1,25,25,0,nfish=,207,,,,,,,,,,,,
3,3,10,13,15,24,17,14,10,16,19,12,20,7,6,7,4,2,0,1,0,1,1,1,1
93,1,5,4,0,220,1,1,25,25,0,nfish=,1231,,,,,,,,,,,,
1,1,3,19,33,38,30,45,50,58,52,78,120,155,122,87,58,57,65,63,29,25,29,12,1
94,1,1,4,3,97,1,1,25,25,0,nfish=,1085,nsamps=,45,,,,,,,,,,
0,0,0,0,0,0,216,1272,3775,2666,3645,10610,6489,8322,12092,4811,4392,560,2774,2012,932,1003,2104,307,0
0,0,0,0,0,108,270,2093,2490,8751,15388,17606,18755,7807,9129,7837,916,1612,3800,946,534,797,0,0,0
94,1,2,4,3,41,1,1,25,25,0,nfish=,516,nsamps=,31,,,,,,,,,,
0,0,0,0,0,0,0,0,86,139,516,814,419,1280,2171,1035,1243,439,203,473,51,0,160,0,0
0,0,0,0,0,0,0,43,99,149,479,761,1086,657,596,290,179,296,127,40,40,40,0,0,0
94,1,3,4,3,54,1,1,25,25,0,nfish=,802,nsamps=,41,,,,,,,,,,
0,0,0,0,0,0,0,0,28,347,937,2668,7577,8562,7744,2641,0,228,577,347,694,394,0,0,0
0,0,0,0,0,0,0,572,31,916,4385,5181,3798,3816,1028,1228,349,0,0,0,0,0,0,0,0
94,1,4,4,0,85,1,1,25,25,0,nfish=,377,,,,,,,,,,,,
0,0,1,0,5,5,20,18,27,29,24,23,31,43,48,29,30,17,5,6,9,4,2,1,0
94,1,5,4,0,139,1,1,25,25,0,nfish=,776,,,,,,,,,,,,
13,15,8,9,6,14,16,47,62,65,62,76,61,65,54,60,39,27,17,17,15,9,13,5,1
95,1,1,4,3,89,1,1,25,25,0,nfish=,675,nsamps=,34,,,,,,,,,,
0,0,0,0,0,0,0,0,1135,722,1245,1596,4843,2575,3809,3385,2474,2890,4301,6036,1052,6006,1194,2006,0
0,0,116,0,0,264,0,1016,35,2092,2030,7474,6464,9455,8122,5351,3407,4898,5547,4561,261,856,1013,0,0
95,1,2,4,3,12,1,1,25,25,0,nfish=,162,nsamps=,9,,,,,,,,,,
0,0,0,0,0,0,0,0,0,15,573,110,472,1152,683,1002,1346,555,468,0,0,0,15,0,0
0,0,0,0,0,101,0,0,0,60,117,603,694,411,72,0,0,173,0,72,0,0,0,0,0

59
95,1,3,4,3,43,1,1,25,25,0,nfish=,563,nsamps=,28,,,,,,,,,,
0,0,0,0,0,0,0,0,64,64,530,905,859,3754,3693,2751,993,407,992,573,516,121,274,0,0
0,0,0,0,0,0,0,0,121,450,2698,4606,4915,2009,753,370,272,0,0,0,0,0,0,0,0
95,1,4,4,0,35,1,1,25,25,0,nfish=,35,,,,,,,,,,,,
0,0,1,0,0,1,0,1,0,3,4,4,3,6,3,4,2,2,0,1,0,0,0,0,0
95,1,5,4,0,145,1,1,25,25,0,nfish=,814,,,,,,,,,,,,
8,2,6,11,44,47,49,34,36,45,70,84,94,62,63,31,35,28,17,15,11,6,8,6,2
95,1,10,4,3,32,1,1,25,25,0,nfish=,,nsamps=,47,,,,,,,,,,
38523,4910,6679,9058,6603,17061,13025,0,0,0,3865,12666,8638,3176,4698,4931,7691,24654,14793,6344,9448,0,7782,4799,0
18880,7366,7366,4911,28832,30902,15926,7303,0,7920,4931,2987,19661,7880,20389,12992,17781,2290,8263,7833,0,0,0,0,0
96,1,1,4,3,88,1,1,25,25,0,nfish=,636,nsamps=,31,,,,,,,,,,
0,0,0,0,0,200,1557,59,2531,8096,6126,2320,1439,3015,4075,8648,3891,2903,3482,1048,87,259,1312,0,35
0,0,0,131,0,0,1688,105,341,7051,6188,2919,14589,6376,4675,760,5227,3663,315,1001,35,0,0,0,0
96,1,2,4,3,41,1,1,25,25,0,nfish=,622,nsamps=,31,,,,,,,,,,
0,0,0,0,0,20,6,0,89,271,312,322,904,1020,1078,920,767,702,271,144,133,0,17,93,0
0,0,0,0,0,47,40,47,182,278,301,674,789,542,443,104,54,0,19,0,0,0,0,0,0
96,1,3,4,3,24,1,1,25,25,0,nfish=,170,nsamps=,7,,,,,,,,,,
0,0,0,0,0,0,175,0,0,0,0,391,1672,2231,4115,2257,1404,190,499,0,0,0,0,0,0
0,0,0,0,0,0,0,0,0,0,648,3009,3642,3270,1821,216,283,190,0,0,0,0,0,0,0
96,1,4,4,0,84,1,1,25,25,0,nfish=,114,,,,,,,,,,,,
1,2,2,6,7,7,6,4,2,3,10,5,10,12,7,13,9,3,1,1,0,1,2,0,0
96,1,5,4,0,146,1,1,25,25,0,nfish=,817,,,,,,,,,,,,
5,2,11,19,44,31,37,35,49,61,56,57,71,67,46,52,47,31,30,22,20,8,10,3,3
97,1,1,4,3,95,1,1,25,25,0,nfish=,991,nsamps=,45,,,,,,,,,,
0,0,0,0,0,0,0,124,129,597,176,472,630,1778,1975,1777,1909,1240,1863,1414,1570,865,875,2688,435
0,0,0,0,0,39,39,94,473,1018,546,2160,2263,3327,1664,2771,4061,2889,2359,613,2062,93,171,0,0
97,1,2,4,3,29,1,1,25,25,0,nfish=,465,nsamps=,22,,,,,,,,,,
0,0,0,0,0,0,0,0,0,41,39,131,189,282,445,569,642,219,473,214,168,155,478,15,0
0,0,0,0,0,0,15,0,0,13,57,87,209,201,84,233,27,15,0,0,0,0,0,0,0
97,1,3,4,3,21,1,1,25,25,0,nfish=,104,nsamps=,4,,,,,,,,,,
0,0,0,0,0,0,0,0,41,0,67,67,391,108,695,497,323,189,175,74,0,0,0,0,0
0,0,0,0,0,0,0,0,41,67,108,190,985,486,263,67,115,74,0,0,0,0,0,0,0
97,1,4,4,0,54,1,1,25,25,0,nfish=,54,,,,,,,,,,,,
0,1,0,1,5,2,2,3,7,11,6,1,4,4,2,2,0,0,2,0,1,0,0,0,0
97,1,5,4,0,314,1,1,25,25,0,nfish=,1759,,,,,,,,,,,,
14,2,5,19,30,42,68,66,91,88,109,114,128,145,137,126,132,145,78,85,47,20,51,15,2
98,1,1,4,3,84,1,1,25,25,0,nfish=,430,nsamps=,24,,,,,,,,,,
0,0,0,0,246,380,213,211,201,164,628,988,1219,1977,1339,699,524,280,1153,702,1265,1170,917,189,27
0,0,0,0,47,344,477,170,549,611,1158,1340,1989,2673,1004,1533,1434,1871,1057,362,299,146,0,0,0
98,1,2,4,3,21,1,1,25,25,0,nfish=,347,nsamps=,16,,,,,,,,,,
0,0,0,0,0,0,15,15,17,91,32,316,229,323,334,416,262,197,49,0,0,40,0,17,0
0,0,0,0,0,0,0,0,51,81,185,257,212,189,183,62,59,0,25,0,0,0,0,0,0
98,1,3,4,3,26,1,1,25,25,0,nfish=,212,nsamps=,10,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,16,819,811,2487,984,154,2387,51,46,35,0,21,0,0,0
0,0,0,0,0,0,0,0,16,42,70,1544,178,216,870,60,0,0,0,8,0,0,0,0,0
98,1,4,4,0,84,1,1,25,25,0,nfish=,106,,,,,,,,,,,,
0,1,0,1,7,9,8,8,11,6,10,13,9,5,4,5,5,2,2,0,0,0,0,0,0
98,1,5,4,0,167,1,1,25,25,0,nfish=,937,,,,,,,,,,,,
10,4,6,31,38,36,13,36,49,72,77,79,79,52,81,59,60,43,37,27,16,14,5,12,1
98,1,10,4,3,25,1,1,25,25,0,nfish=,,nsamps=,37,,,,,,,,,,
2284,0,0,7550,21844,21579,10836,6785,0,0,0,3677,0,0,1734,0,0,3612,0,2098,2788,0,3113,0,0
0,0,4407,2997,12517,5118,2627,0,0,0,0,1366,2178,13373,3612,0,2178,5118,0,3128,0,0,0,0,0
99,1,1,4,3,84,1,1,25,25,0,nfish=,424,nsamps=,17,,,,,,,,,,
0,0,0,0,0,0,398,1242,1755,1006,240,111,272,465,432,212,232,221,484,943,150,70,460,714,0
0,0,0,21,19,324,570,1168,1074,426,287,277,473,767,907,662,967,599,450,399,0,0,0,0,0
99,1,2,4,3,8,1,1,25,25,0,nfish=,114,nsamps=,6,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,148,0,218,256,228,832,660,328,140,298,164,0,0,0,0
0,0,0,0,0,0,0,74,0,0,74,74,186,636,446,238,490,350,612,112,112,0,112,0,0
99,1,4,4,0,85,1,1,25,25,0,nfish=,421,,,,,,,,,,,,
0,3,4,1,3,4,11,7,13,15,28,34,46,63,40,45,41,21,22,11,2,2,3,2,0
99,1,5,4,0,114,1,1,25,25,0,nfish=,637,,,,,,,,,,,,
2,0,1,3,10,13,11,38,44,47,34,41,57,71,47,40,50,30,35,21,22,8,6,6,0
100,1,1,4,3,80,1,1,25,25,0,nfish=,191,nsamps=,10,,,,,,,,,,
0,0,0,26,158,39,39,79,0,159,276,458,249,432,273,0,101,148,77,53,112,150,56,37,48
0,0,0,105,39,92,105,79,106,88,271,572,267,232,53,19,114,55,20,48,27,21,0,0,0
100,1,2,4,3,12,1,1,25,25,0,nfish=,69,nsamps=,9,,,,,,,,,,

60
0,0,0,28,0,0,0,0,0,0,66,28,0,160,94,132,56,132,28,0,0,0,28,0,0
0,0,0,0,0,0,0,0,0,66,160,226,122,132,160,28,28,56,56,56,56,28,0,0,0
100,1,4,4,0,85,1,1,25,25,0,nfish=,505,,,,,,,,,,,,
30,69,85,40,31,14,7,2,5,8,12,8,27,20,26,35,27,18,20,11,6,4,0,0,0
100,1,5,4,0,50,1,1,25,25,0,nfish=,282,,,,,,,,,,,,
10,26,19,12,8,18,10,14,15,18,26,7,15,9,4,5,6,5,11,8,17,10,5,1,3
101,1,1,4,3,88,1,1,25,25,0,nfish=,617,nsamps=,25,,,,,,,,,,
0,0,3,60,228,235,144,229,211,82,29,50,91,121,147,44,75,6,123,17,22,17,167,0,77
0,0,12,75,247,255,103,235,49,31,54,91,250,131,42,25,6,92,474,0,158,0,77,0,0
101,1,2,4,3,24,1,1,25,25,0,nfish=,233,nsamps=,18,,,,,,,,,,
0,0,0,0,0,18,90,56,9,20,20,42,49,31,76,114,90,147,51,62,81,34,25,0,0
0,0,0,3,9,18,45,57,20,0,33,69,51,88,91,84,82,43,80,46,0,0,0,0,0
101,1,4,4,0,85,1,1,25,25,0,nfish=,380,,,,,,,,,,,,
1,1,10,27,80,78,78,38,12,10,4,3,9,9,7,1,4,2,2,2,2,0,0,0,0
101,1,5,4,0,58,1,1,25,25,0,nfish=,324,,,,,,,,,,,,
1,2,1,10,27,53,48,17,16,15,11,23,17,17,24,10,12,10,4,1,1,0,2,2,0
101,1,10,4,3,21,1,1,25,25,0,nfish=,114,nsamps=,31,,,,,,,,,,
0,0,0,2367,28385,11560,25465,3103,5351,2289,2289,0,0,0,2516,0,0,2516,0,0,0,0,2744,0,0
0,0,0,2539,21189,22364,34500,2984,5031,0,0,0,0,8417,3103,4878,5804,0,2047,2047,0,0,0,0,0
102,1,1,4,3,82,1,1,25,25,0,nfish=,320,nsamps=,15,,,,,,,,,,
0,0,0,0,0,0,0,218,510,552,341,337,123,54,236,393,114,173,163,153,340,131,120,0,70
0,0,0,0,57,78,93,259,661,307,281,199,178,336,61,73,0,0,90,30,3,0,0,0,0
102,1,2,4,3,1,1,1,25,25,0,nfish=,14,nsamps=,1,,,,,,,,,,
0,0,0,0,0,0,0,0,0,2,2,0,2,4,8,2,2,0,0,0,0,0,0,0,0
0,0,0,0,0,0,0,0,0,0,0,0,0,0,6,0,0,0,0,0,0,0,0,0,0
102,1,3,4,3,17,1,1,25,25,0,nfish=,25,nsamps=,1,,,,,,,,,,
0,0,0,0,0,0,0,0,0,0,0,0,0,2,0,0,0,0,0,0,0,0,0,2,0
0,0,0,0,0,0,0,0,0,0,0,2,2,10,20,14,0,0,0,0,0,0,0,0,0
102,1,4,4,0,86,1,1,25,25,0,nfish=,771,,,,,,,,,,,,
0,1,2,3,2,20,39,83,137,139,117,72,22,21,31,27,16,13,15,4,3,2,2,0,0
102,1,5,4,0,32,1,1,25,25,0,nfish=,180,,,,,,,,,,,,
0,0,0,0,0,1,1,7,29,43,33,17,2,8,7,11,6,8,3,2,2,0,0,0,0
103,1,4,4,0,84,1,1,25,25,0,nfish=,122,,,,,,,,,,,,
0,2,0,0,0,0,0,2,14,16,21,29,17,4,5,6,0,3,1,1,1,0,0,0,0
104,1,1,4,0,78,1,1,25,25,0,nfish=,110,nsamps=,9,,,,,,,,,,
0,0,0,0,0,0,0,25,4,29,33,80,65,73,288,168,181,60,44,0,57,0,16,0,0
104,1,4,4,0,86,1,1,25,25,0,nfish=,889,,,,,,,,,,,,
8,17,27,44,24,27,20,25,48,55,105,135,116,97,52,37,21,8,8,5,4,2,2,0,2
104,1,5,4,0,14,1,1,25,25,0,nfish=,80,,,,,,,,,,,,
0,0,1,0,2,1,3,2,9,6,5,9,4,9,4,8,2,6,1,2,2,1,3,0,0
104,1,10,4,3,23,1,1,25,25,0,nfish=,216,nsamps=,33,,,,,,,,,,
19065,29801,2189,0,0,0,0,0,0,42045,2356,26204,56703,59820,99071,121898,48127,28657,19776,19405,21625,11473,3,7044,4580
13679,29672,4912,2456,0,0,2456,0,0,45264,9674,14121,23379,115715,61595,72185,65801,27955,31884,16194,35618,15942,0,0,0
105,1,4,4,0,84,1,1,25,25,0,nfish=,571,,,,,,,,,,,,
0,0,2,10,20,18,3,2,4,9,9,11,8,13,12,8,6,0,1,0,1,0,0,0,0
105,1,5,4,0,13,1,1,25,25,0,nfish=,73,,,,,,,,,,,,
0,1,0,1,1,2,0,2,1,0,4,5,8,14,13,10,3,4,1,1,2,0,0,0,0
106,1,2,4,0,13,1,1,25,25,0,nfish=,78,nsamps=,10,,,,,,,,,,
0,0,0,0,0,20,20,40,175,72,111,119,119,69,58,153,48,121,93,14,23,35,45,0,0
106,1,4,4,0,88,1,1,25,25,0,nfish=,1330,,,,,,,,,,,,
2,3,6,10,29,54,101,183,265,189,89,54,55,54,78,66,43,26,9,4,2,4,2,1,1
106,1,5,4,0,9,1,1,25,25,0,nfish=,49,,,,,,,,,,,,
0,0,0,0,0,1,0,5,4,1,3,4,3,5,8,4,6,3,1,0,0,0,1,0,0

61
Appendix C. Parameter file for model STATc
boc2007revC.csv **** UNKNOWN CONVERGENCE STATUS
statc07revC.r01
statc07revC.par
2007 starting from statc2005 assessment postSTAR include all & 0.1srr, revised c
10.000000 .000100 BEGIN AND END DELTA F PER LOOP1
3 .95 FIRST LOOP1 FOR LAMBDA & VALUE
1.100 MAX VALUE FOR CROSS DERIVATIVE
1 READ HESSIAN
STARB2.hes
1 WRITE HESSIAN
STARB2.hes
.001 MIN SAMPLE FRAC. PER AGE
1 21 1 21 MINAGE, MAXAGE, SUMMARY AGE RANGE
51 106 BEGIN YEAR, END YEAR
1 12 0 0 0 NPER, MON/PER
1.00 SPAWNMONTH
5 9 NFISHERY, NSURVEY
2 N SEXES
50000. REF RECR LEVEL
0 MORTOPT
.150000 .010000 .250000 'M ' 0 1 0 .000000 .0000 ! 1 NO PICK .000 0. .0000000
-999.000000 .010000 1.000000 'M SAME FOR M+F ' 0 1 0 .000000 .0000 ! 2 NO PICK .000 0. .0000000
TRAWL TYPE: 1
7 SELECTIVITY PATTERN
0 0 0 2 0 0 0 AGE TYPES USED
1.00000 .10 ' TWL CATCH BIOMASS ' ! # = 1 VALUE: .00000
1.00000 .30 ' TWL SIZE COMPS ' ! # = 2 VALUE: -537.43774
1 1 0 0 0 0 SEL. COMPONENTS
51.916183 20.000000 70.000000 'Trawl:transition' 2 1 0 .000000 .0000 ! 3 OK -.005 -27. 1672.6971612
.000001 .000001 1.000000 'Trawl:InitSelect' 0 1 0 .000000 .0000 ! 4 NO PICK .000 0. .0000000
.488685 .001000 1.000000 'Trawl:SmlInflect' 2 1 0 .500000 1.0000 ! 5 OK .000 -45676. .5484042
.336529 .001000 3.000000 'Trawl:SmlSlope ' 2 1 0 .900000 1.0000 ! 6 OK .000 -3199. .0015743
.595213 .001000 1.000000 'Trawl:femfinal ' 2 1 0 1.000000 1.0000 ! 7 OK .000 -4823. .0066239
.348224 .001000 1.000000 'Trawl:feminflct ' 2 1 0 .500000 1.0000 ! 8 OK .000 -2896. 1.5064589
1.347485 .001000 5.000000 'Trawl:femSlope ' 0 1 0 .900000 1.0000 ! 9 NO PICK .000 0. .0000000
H&L TYPE: 2
7 SELECTIVITY PATTERN
0 0 0 4 0 0 0 AGE TYPES USED
1.00000 .10 ' H&Lso CATCH BIOMASS' ! # = 3 VALUE: .00000
1.00000 .30 ' H&Lso SIZE COMPS ' ! # = 4 VALUE: -204.38755
1 1 0 0 0 0 SEL. COMPONENTS
48.382066 20.000000 70.000000 'H&L:transition ' 2 1 0 .000000 .0000 ! 10 OK .000 -7. 5.2163052
.003059 .000001 1.000000 'H&L:InitSelect ' 2 1 0 .000000 .0000 ! 11 OK .000 -56790. .0001048
.840932 .001000 1.000000 'H&L:SmlInflect ' 2 1 0 .500000 1.0000 ! 12 OK .000 -1877. .0306416
.333099 .001000 3.000000 'H&L:SmlSlope ' 2 1 0 .900000 1.0000 ! 13 OK .000 -2281. .0114772
.275881 .001000 1.000000 'H&L:femfinal ' 2 1 0 1.000000 1.0000 ! 14 OK .000 -363. .0798732
.380517 .001000 1.000000 'H&L:feminflct ' 2 1 0 .500000 1.0000 ! 15 OK .000 -305. .0222269
.268922 .001000 5.000000 'H&L:femSlope ' 2 1 0 .900000 1.0000 ! 16 OK .000 -89. .1777621
SETNET TYPE: 3
7 SELECTIVITY PATTERN
0 0 0 6 0 0 0 AGE TYPES USED
1.00000 .10 'SetNetCATCHBIOM ' ! # = 5 VALUE: .00000
1.00000 .30 'SetNetSizeComps ' ! # = 6 VALUE: -258.61169
1 1 0 0 0 0 SEL. COMPONENTS
49.540604 20.000000 60.000000 'StNso:transition' 2 1 0 .000000 .0000 ! 17 OK .004 -19. 23.4146939
.004154 .000001 1.000000 'StNso:InitSelect' 2 1 0 .000000 .0000 ! 18 OK .000 -313878. .0000070
.785004 .001000 .990000 'StNso:YngInflect' 2 1 0 .500000 1.0000 ! 19 OK .000 -11461. .0296508
.653476 .001000 3.000000 'StNso:YngSlope ' 2 1 0 .900000 1.0000 ! 20 OK .000 -458. .0135755
.147646 .001000 1.000000 'StNso:femfinal ' 2 1 0 .000000 .0000 ! 21 OK .000 -1213. .0059668
.131565 .001000 1.000000 'StNso:feminflct ' 2 1 0 .500000 1.0000 ! 22 OK -.001 -344. .3510564
.247784 .001000 5.000000 'StNso:femSlope ' 2 1 0 .900000 1.0000 ! 23 OK .000 -1044. .0459814
RECLso TYPE: 4
7 SELECTIVITY PATTERN

62
0 0 0 8 0 0 0 AGE TYPES USED
1.00000 .10 'RECLsoCATCHBIOM ' ! # = 7 VALUE: .00000
1.00000 .30 'RECLsoSIZECOMPS ' ! # = 8 VALUE: -310.73584
1 1 0 0 0 0 SEL. COMPONENTS
41.756833 15.000000 60.000000 'RCLso:transition' 2 1 0 .000000 .0000 ! 24 OK -.003 -3. 53.1311282
.140186 .000001 1.000000 'RCLso:InitSelect' 2 1 0 .000000 .0000 ! 25 OK .000 -2921. .0004495
.001000 .001000 1.000000 'RCLso:SmlInflect' 0 1 0 .500000 1.0000 ! 26 NO PICK .000 0. .0000000
.194675 .001000 5.000000 'RCLso:SmlSlope ' 2 1 0 .900000 1.0000 ! 27 OK .000 -1909. .0027112
.067805 .001000 1.000000 'RCLso:femfinal ' 2 1 0 .000000 .0000 ! 28 OK .000 -4034. .0012391
.386057 .001000 1.000000 'RCLso:feminflct ' 2 1 0 .500000 1.0000 ! 29 OK .000 -5940. .0252360
.298196 .001000 5.000000 'RCLso:femSlope ' 2 1 0 .900000 1.0000 ! 30 OK .000 -941. .0045968
RECLnor TYPE: 5
7 SELECTIVITY PATTERN
0 0 0 10 0 0 0 AGE TYPES USED
1.00000 .10 'RECLnorCATCHBIOM ' ! # = 9 VALUE: .00000
1.00000 .30 'RECLnorSIZECOMPS ' ! # = 10 VALUE: -267.58865
1 1 0 0 0 0 SEL. COMPONENTS
48.733482 15.000000 60.000000 'RCLno:transition' 2 1 0 .000000 .0000 ! 31 OK -.001 -2. 164.7240616
.065319 .000001 1.000000 'RCLno:InitSelect' 2 1 0 .000000 .0000 ! 32 OK .000 -12248. .0014999
.496223 .001000 1.000000 'RCLno:SmlInflect' 2 1 0 .500000 1.0000 ! 33 OK .000 -606. .6273304
.129680 .001000 5.000000 'RCLno:SmlSlope ' 2 1 0 .900000 1.0000 ! 34 OK .000 -1046. .2221516
.279615 .001000 1.000000 'RCLno:femfinal ' 2 1 0 .000000 .0000 ! 35 OK .000 -398. .0133749
.584202 .001000 1.000000 'RCLno:feminflct ' 2 1 0 .500000 1.0000 ! 36 OK .000 -686. .0526584
.268031 .001000 5.000000 'RCLno:femSlope ' 2 1 0 .900000 1.0000 ! 37 OK .000 -114. .0271967
NoRec TYPE: 6
2 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.000168 0 1 2 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
1.00000 .67 'RecFINnoCPUE ' ! # = 11 VALUE: -4.91890
5.000000 -.200000 1.000000 'NoCalCPU:Seltype' 0 -80 0 .000000 .0000 ! 38 NO PICK .000 0. .0000000
24.000000 .010000 24.000000 'NoCalCPUl:minsiz' 0 -80 0 .000000 .0000 ! 39 NO PICK .000 0. .0000000
76.000000 .001000 76.000000 'NoCalCPUl:maxsiz' 0 -80 0 .000000 .0000 ! 40 NO PICK .000 0. .0000000
DFGcpuN TYPE: 7
2 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.000733 0 1 2 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
1.00000 .37 ' NoCalDFG ' ! # = 12 VALUE: 8.19614
5.000000 -.200000 1.000000 'NoCalDFG:Seltyp ' 0 -87 0 .000000 .0000 ! 41 NO PICK .000 0. .0000000
24.000000 .010000 24.000000 'NoCalDFG:minsi ' 0 -87 0 .000000 .0000 ! 42 NO PICK .000 0. .0000000
76.000000 .001000 76.000000 'NOCalDFG:maxsi ' 0 -87 0 .000000 .0000 ! 43 NO PICK .000 0. .0000000
SoRecFI TYPE: 8
2 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.000196 0 1 2 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
1.00000 .71 'RecFINsoCPUE ' ! # = 13 VALUE: -4.06754
4.000000 -.200000 1.000000 'SoCalCPU:Seltype' 0 -80 0 .000000 .0000 ! 44 NO PICK .000 0. .0000000
24.000000 .010000 24.000000 'SoCalCPUl:minsiz' 0 -80 0 .000000 .0000 ! 45 NO PICK .000 0. .0000000
76.000000 .001000 76.000000 'SoCalCPUl:maxsiz' 0 -80 0 .000000 .0000 ! 46 NO PICK .000 0. .0000000
TwlCPUE TYPE: 9
2 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.004940 0 1 1 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
1.00000 .32 'TrawlCPUE ' ! # = 14 VALUE: 9.45784
1.000000 -.200000 1.000000 'TrawlSeltype ' 0 -82 0 .000000 .0000 ! 47 NO PICK .000 0. .0000000
20.000000 .010000 20.000000 'TrawlCPUE:minsiz' 0 -82 0 .000000 .0000 ! 48 NO PICK .000 0. .0000000
84.000000 .001000 84.000000 'TrawlCPUE:maxsiz' 0 -82 0 .000000 .0000 ! 49 NO PICK .000 0. .0000000
TRITRAW TYPE: 10
7 SELECTIVITY PATTERN
0 0 0 16 0 0 0 AGE TYPES USED
.044151 0 1 1 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
1.00000 .81 'TRI SURVEY BIO ' ! # = 15 VALUE: -5.56087
1.00000 .30 'TRI SIZE COMPS ' ! # = 16 VALUE: -82.35644
1 1 0 0 0 0 SEL. COMPONENTS
34.311827 26.000000 76.000000 'TriSv:transition' 0 89 0 .000000 .0000 ! 50 NO PICK .000 0. .0000000
.363817 .001000 1.000000 'TriSv:InitSelect' 2 89 0 .000000 .0000 ! 51 OK .000 -55. .0195570
.001000 .001000 1.000000 'TriSv:YngInflect' 0 89 0 .500000 1.0000 ! 52 NO PICK .000 0. .0000000

63
 3.000000 .001000 3.000000 'TriSv:YngSlope ' 0 89 0 .900000 1.0000 ! 53 NO PICK .000 0. .0000000
.484561 .001000 1.000000 'TriSv:femfinal ' 2 89 0 .000000 .0000 ! 54 OK .000 -169. .0127043
.001000 .001000 1.000000 'TriSv:feminflct ' 0 89 0 .500000 1.0000 ! 55 NO PICK .000 0. .0000000
5.000000 .001000 5.000000 'TriSv:femSlope ' 0 89 0 .900000 1.0000 ! 56 NO PICK .000 0. .0000000
CALCOFI TYPE: 11
4 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.000233 0 1 1 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
1.00000 .68 'CALCOFISPB ' ! # = 17 VALUE: -1.11389
PowPlnt TYPE: 12
3 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.011220 0 1 2 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
.00000 2.10 'PowPltRectIndex ' ! # = 18 VALUE: -35.54367
1.000000 .000000 1.000000 'PowplntAge1Nos ' 0 -73 0 .000000 .0000 ! 57 NO PICK .000 0. .0000000
1.000000 .000000 1.000000 'PowplntAge1Nos ' 0 -73 0 .000000 .0000 ! 58 NO PICK .000 0. .0000000
JuvSurv TYPE: 13
3 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.000078 0 1 2 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
.00000 2.05 'CenCalJuvIndex ' ! # = 19 VALUE: -25.19993
1.000000 .000000 1.000000 'JuvSurvAge1Nos ' 0 -84 0 .000000 .0000 ! 59 NO PICK .000 0. .0000000
1.000000 .000000 1.000000 'JuvSurvAge1Nos ' 0 -84 0 .000000 .0000 ! 60 NO PICK .000 0. .0000000
PierCPU TYPE: 14
3 SELECTIVITY PATTERN
0 0 0 0 0 0 0 AGE TYPES USED
.000275 0 1 2 Q, QUANT, LOGERROR=1, BIO=1 or NUM=2
.00000 3.29 'PierRectIndex ' ! # = 20 VALUE: -32.84073
1.000000 .000000 1.000000 'PierIndex1Nos ' 0 -81 0 .000000 .0000 ! 61 NO PICK .000 0. .0000000
1.000000 .000000 1.000000 'PierIndex1Nos ' 0 -81 0 .000000 .0000 ! 62 NO PICK .000 0. .0000000
1 AGEERR: 1: MULTINOMIAL, 0: S(LOG(P))=CONSTANT, -1: S=P*Q/N
500.000 : MAX N FOR MULTINOMIAL
3 1=%CORRECT, 2=C.V., 3=%AGREE, 4=READ %AGREE @AGE
.800000 .300000 .950000 'p AGREE. @1 ' 0 80 0 .000000 .0000 ! 63 NO PICK .000 0. .0000000
.050000 .000000 .900000 'p agree @21 ' 0 80 0 .000000 .0000 ! 64 NO PICK .000 0. .0000000
1.000000 .001000 2.000000 'POWER ' 0 80 0 .000000 .0000 ! 65 NO PICK .000 0. .0000000
.150000 .010000 .300000 'OLD DISCOUNT ' 0 80 0 .000000 .0000 ! 66 NO PICK .000 0. .0000000
.000001 .001000 .100000 '%MIS-SEXED ' 0 80 0 .000000 .0000 ! 67 NO PICK .000 0. .0000000
0 END OF EFFORT
0 FIX n FMORTs
0 MATURITY
1 GROWTH: 1=CONSTANT, 2=MORT. INFLUENCE
1.5000 99.0000 AGE AT WHICH L1 AND L2 OCCUR
1 1=NORMAL, 2=LOGNORMAL
27.000000 20.000000 60.000000 'FEMALE L1 ' 0 1 0 .000000 .0000 ! 68 NO PICK .000 0. .0000000
75.892728 60.000000 90.000000 'FEMALE LINF ' 0 1 0 .000000 .0000 ! 69 NO PICK .000 0. .0000000
.183673 .050000 .400000 'FEMALE K ' 2 1 0 .000000 .0000 ! 70 OK .000-2186705. .0000036
.107000 .010000 .990000 'FEMALE CV1 ' 0 1 0 .000000 .0000 ! 71 NO PICK .000 0. .0000000
.033000 .010000 .990000 'FEMALE CV21 ' 0 1 0 .000000 .0000 ! 72 NO PICK .000 0. .0000000
-999.000000 20.000000 40.000000 'MALE L1 ' 0 1 0 .000000 .0000 ! 73 NO PICK .000 0. .0000000
65.555310 50.000000 80.000000 'MALE LINF ' 0 1 0 .000000 .0000 ! 74 NO PICK .000 0. .0000000
.210373 .100000 .400000 'MALE K ' 2 1 0 .000000 .0000 ! 75 OK .000-1146616. .0001222
-999.000000 .010000 .990000 'MALE CV1 ' 0 1 0 .000000 .0000 ! 76 NO PICK .000 0. .0000000
-999.000000 .010000 .990000 'MALE CV21 ' 0 1 0 .000000 .0000 ! 77 NO PICK .000 0. .0000000
0 DEFINE MARKET CATEGORIES
0 ENVIRONMENTAL FXN: [-INDEX] [FXN TYPE(1-4)] [ENVVAR USED]
0 ESTIMATE N ENVIRON VALUES
21 PENALTIES
.00000 .30 ' Parm Penalty ' ! # = 21 VALUE: -40.61406
-1 1.0 1.0
0 ENVIRONMENT EFFECT ON EXP(RECR)
22 STOCK-RECR
3 1=B-H, 2=RICKER, 3=new B-H, 4=HOCKEY
0 disabled option
.10000 -1.00 'SPAWN RECR. ' ! # = 22 VALUE: -44.15689
.00001 -.30 'S-R means ' ! # = 23 VALUE: -527.37648

64
 10.000000 .001000 10.000000 'VIR. RECR. MULT.' 0 1 0 .000000 .0000 ! 78 NO PICK .000 0. .0000000
.200000 .200000 .990000 'B-H S/R PAR. ' 0 1 0 .000000 .0000 ! 79 NO PICK .000 0. .0000000
.070451 .001000 10.000000 'BACK RECR. ' 0 1 0 .000000 .0000 ! 80 NO PICK .000 0. .0000000
1.000000 .010000 2.000000 'S/R STD. ' 0 1 0 .000000 .0000 ! 81 NO PICK .000 0. .0000000
.000000 -.100000 .100000 'RECR. TREND ' 0 1 0 .000000 .0000 ! 82 NO PICK .000 0. .0000000
1.000000 .000000 2.000000 'RECR. MULT. ' 0 1 0 .000000 .0000 ! 83 NO PICK .000 0. .0000000
-2 INIT AGE COMP
-999.000000 .001000 30.000000 'Recruit 51 ' -2 51 0 .000000 .0000 ! 84 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 52 ' -2 52 0 .000000 .0000 ! 85 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 53 ' -2 53 0 .000000 .0000 ! 86 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 54 ' -2 54 0 .000000 .0000 ! 87 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 55 ' -2 55 0 .000000 .0000 ! 88 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 56 ' -2 56 0 .000000 .0000 ! 89 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 57 ' -2 57 0 .000000 .0000 ! 90 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 58 ' -2 58 0 .000000 .0000 ! 91 NO PICK .000 0. .0000000
-999.000000 .001000 30.000000 'Recruit 59 ' -2 59 0 .000000 .0000 ! 92 NO PICK .000 0. .0000000
.113969 .001000 30.000000 'Recruit 60 ' 2 60 0 .000000 .0000 ! 93 OK .000 -6425. 2.2671210
.026661 .001000 30.000000 'Recruit 61 ' 2 61 0 .000000 .0000 ! 94 OK .000 -7798. .0332638
.023849 .001000 30.000000 'Recruit 62 ' 2 62 0 .000000 .0000 ! 95 OK .000 -9434. .0281218
1.046731 .001000 30.000000 'Recruit 63 ' 2 63 0 .000000 .0000 ! 96 OK .000 -11018. 1.2380030
.017206 .001000 30.000000 'Recruit 64 ' 2 64 0 .000000 .0000 ! 97 OK .000 -13153. .0115736
.015136 .001000 30.000000 'Recruit 65 ' 2 65 0 .000000 .0000 ! 98 OK .000 -15536. .0040839
.017790 .001000 30.000000 'Recruit 66 ' 2 66 0 .000000 .0000 ! 99 OK .000 -18350. .0064730
.026870 .001000 30.000000 'Recruit 67 ' 2 67 0 .000000 .0000 ! 100 OK .000 -21294. .0415621
.043123 .001000 30.000000 'Recruit 68 ' 2 68 0 .000000 .0000 ! 101 OK .000 -23888. .1460500
.060870 .001000 30.000000 'Recruit 69 ' 2 69 0 .000000 .0000 ! 102 OK .000 -25965. .0289535
.063977 .001000 30.000000 'Recruit 70 ' 2 70 0 .000000 .0000 ! 103 OK .000 -27508. .0330586
.287270 .001000 30.000000 'Recruit 71 ' 2 71 0 .000000 .0000 ! 104 OK .000 -29243. .0873928
.036365 .001000 30.000000 'Recruit 72 ' 2 72 0 .000000 .0000 ! 105 OK .000 -32986. .0280150
.039638 .001000 30.000000 'Recruit 73 ' 2 73 0 .000000 .0000 ! 106 OK .000 -38974. .0061877
.317521 .001000 30.000000 'Recruit 74 ' 2 74 0 .000000 .0000 ! 107 OK .000 -44829. .0009801
.110900 .001000 30.000000 'Recruit 75 ' 2 75 0 .000000 .0000 ! 108 OK .000 -53937. .0003446
.025811 .001000 30.000000 'Recruit 76 ' 2 76 0 .000000 .0000 ! 109 OK .000 -72778. .0000723
.010744 .001000 30.000000 'Recruit 77 ' 2 77 0 .000000 .0000 ! 110 OK .000 -85472. .0000415
.475827 .001000 30.000000 'Recruit 78 ' 2 78 0 .000000 .0000 ! 111 OK .000 -59696. .0007582
.039606 .001000 30.000000 'Recruit 79 ' 2 79 0 .000000 .0000 ! 112 OK .000 -61134. .0006102
.167462 .001000 30.000000 'RECRUIT 80 ' 2 80 0 .000000 .0000 ! 113 OK .000 -65065. .0004003
.027938 .001000 30.000000 'RECRUIT 81 ' 2 81 0 .000000 .0000 ! 114 OK .000 -82601. .0001300
.031985 .001000 30.000000 'RECRUIT 82 ' 2 82 0 .000000 .0000 ! 115 OK .000 -122236. .0000850
.003014 .001000 30.000000 'RECRUIT 83 ' 2 83 0 .000000 .0000 ! 116 OK .000 -297957. .0000076
.012743 .001000 30.000000 'RECRUIT 84 ' 2 84 0 .000000 .0000 ! 117 OK .000 -160645. .0000216
.216766 .001000 30.000000 'RECRUIT 85 ' 2 85 0 .000000 .0000 ! 118 OK .000 -89087. .0001718
.029077 .001000 30.000000 'RECRUIT 86 ' 2 86 0 .000000 .0000 ! 119 OK .000 -122308. .0000765
.028413 .001000 30.000000 'RECRUIT 87 ' 2 87 0 .000000 .0000 ! 120 OK .000 -153583. .0000179
.033653 .001000 30.000000 'RECRUIT 88 ' 2 88 0 .000000 .0000 ! 121 OK .000 -133208. .0000195
.117140 .001000 30.000000 'RECRUIT 89 ' 2 89 0 .000000 .0000 ! 122 OK .000 -92186. .0000478
.002850 .001000 30.000000 'RECRUIT 90 ' 2 90 0 .000000 .0000 ! 123 OK .000 -191316. .0000158
.037624 .001000 30.000000 'RECRUIT 91 ' 2 91 0 .000000 .0000 ! 124 OK .000 -157201. .0000164
.030549 .001000 30.000000 'RECRUIT 92 ' 2 92 0 .000000 .0000 ! 125 OK .000 -195647. .0000420
.007710 .001000 30.000000 'RECRUIT 93 ' 2 93 0 .000000 .0000 ! 126 OK .000 -337838. .0000195
.017371 .001000 30.000000 'RECRUIT 94 ' 2 94 0 .000000 .0000 ! 127 OK .000 -347802. .0000125
.015919 .001000 30.000000 'RECRUIT 95 ' 2 95 0 .000000 .0000 ! 128 OK .000 -392818. .0000073
.008697 .001000 30.000000 'RECRUIT 96 ' 2 96 0 .000000 .0000 ! 129 OK .000 -460757. .0000042
.020112 .001000 30.000000 'RECRUIT 97 ' 2 97 0 .000000 .0000 ! 130 OK .000 -311071. .0000131
.004891 .001000 30.000000 'RECRUIT 98 ' 2 98 0 .000000 .0000 ! 131 OK .000 -482180. .0000058
.007352 .001000 30.000000 'RECRUIT 99 ' 2 99 0 .000000 .0000 ! 132 OK .000 -205674. .0000094
.118887 .001000 30.000000 'RECRUIT 100 ' 2 100 0 .000000 .0000 ! 133 OK .000 -13098. .0004233
.001000 .001000 30.000000 'RECRUIT 101 ' 0 101 0 .000000 .0000 ! 134 NO PICK .000 0. .0000000
.009630 .001000 30.000000 'RECRUIT 102 ' 2 102 0 .000000 .0000 ! 135 OK .000 -99226. .0000237
.009787 .001000 30.000000 'RECRUIT 103 ' 2 103 0 .000000 .0000 ! 136 OK .000 -82446. .0000204
.054638 .001000 30.000000 'RECRUIT 104 ' 2 104 0 .000000 .0000 ! 137 OK .000 -16128. .0001699
-.018345 .001000 30.000000 'RECRUIT 105 ' -2 105 0 .000000 .0000 ! 138 NO PICK .000 0. .0000000
-.020981 .001000 30.000000 'RECRUIT 106 ' -2 106 0 .000000 .0000 ! 139 NO PICK .000 0. .0000000

65
Appendix E. Summary of results.
Table E1. Results of bocaccio model STATc2007. Bage1+ is summary biomass of age 1 and older fish (mtons). SpOut is spawning
output in 10E9eggs. Recruits is of age 1 fish (10E3 fish). Catch is in mtons. ExploitRate is exploitation rate (Catch/Bage1+).
Year Bage1+ SpOut Recruits Catch ExploitRate Year Bage1+ SpOut Recruits Catch ExploitRate
pre-1951 22625 3580 3523 2000 8.8% 1978 28130 4027 23791 3565 12.7%
1951 22625 3580 3523 2148 9.5% 1979 28586 3923 1980 4878 17.1%
1952 22478 3560 3523 2098 9.3% 1980 29561 3600 8373 6125 20.7%
1953 22381 3547 3523 2370 10.6% 1981 27814 3552 1397 5680 20.4%
1954 22007 3486 3523 2529 11.5% 1982 25511 3584 1599 6518 25.5%
1955 21468 3396 3523 2645 12.3% 1983 20837 3305 151 5587 26.8%
1956 20807 3285 3523 3116 15.0% 1984 16131 2837 637 4671 29.0%
1957 19667 3088 3523 3207 16.3% 1985 13504 2256 10838 2870 21.3%
1958 18457 2858 3523 3558 19.3% 1986 12129 1858 1454 3123 25.7%
1959 16936 2565 3523 3018 17.8% 1987 11034 1459 1421 2671 24.2%
1960 16405 2359 5698 2702 16.5% 1988 10324 1351 1683 2324 22.5%
1961 15787 2221 1333 2151 13.6% 1989 10291 1349 5857 2773 26.9%
1962 15597 2213 1192 1836 11.8% 1990 9039 1179 143 2758 30.5%
1963 24259 2311 52337 2503 10.3% 1991 7946 968 1881 1782 22.4%
1964 29563 2312 860 1667 5.6% 1992 7552 958 1527 1874 24.8%
1965 38092 2801 757 1971 5.2% 1993 6693 921 385 1565 23.4%
1966 44622 4331 890 3258 7.3% 1994 5948 857 869 1216 20.4%
1967 47280 6221 1344 5003 10.6% 1995 5348 820 796 774 14.5%
1968 45736 7227 2156 3029 6.6% 1996 5037 808 435 566 11.2%
1969 44532 7736 3044 2106 4.7% 1997 4944 804 1006 479 9.7%
1970 43288 7910 3199 2451 5.7% 1998 4796 802 245 206 4.3%
1971 43158 7766 14364 2459 5.7% 1999 4888 836 368 197 4.0%
1972 42115 7499 1818 3623 8.6% 2000 5882 871 5944 189 3.2%
1973 40308 7063 1982 7734 19.2% 2001 6522 901 50 207 3.2%
1974 36304 6137 15876 8494 23.4% 2002 7422 958 481 132 1.8%
1975 30969 5034 5545 5750 18.6% 2003 8213 1134 489 14 0.2%
1976 28518 4335 1291 5410 19.0% 2004 9283 1386 2732 82 0.9%
1977 25775 3961 537 3348 13.0% 2005 10024 1585 917 87 0.9%
2006 10752 1727 1049 67 0.6%

66
Table E2. Age structure in 2006. BEGWT is weight (kg)at the beginning of the year. MIDWT is weight (kg) in the middle of the year.
MATURE is fraction mature. EGGS is eggs per gram for mature fish. SPAWN is net fucundity at age. N106 is abundance (10E3 fish)
at age at the beginningof 2006. RELATIVE F is fishing mortality rate relative to the maximum value.

FEMALES MALES
AGE BEGWT MIDWT MATURE EGGS SPAWN N 106 RELATIVE F BEGWT MIDWT N 106 RELATIVE F
1 0.1741 0.2227 0.0018 0.232 0.0001 524.5 0.21 0.1766 0.2235 524.5 0.21
2 0.3668 0.4983 0.0203 0.2396 0.0018 394 0.56 0.3495 0.4604 393.9 0.52
3 0.7092 0.8752 0.1388 0.2529 0.0257 1002.4 0.81 0.6322 0.7631 1002.8 0.75
4 1.1233 1.3083 0.4165 0.2688 0.1296 153 0.98 0.9528 1.0904 153.2 0.93
5 1.5736 1.7649 0.6992 0.286 0.322 128.1 0.96 1.2823 1.4172 128.2 1.00
6 2.0314 2.2191 0.8667 0.3035 0.5436 11.2 0.82 1.6006 1.7266 11.3 0.98
7 2.476 2.6541 0.9423 0.3204 0.7579 1119.3 0.66 1.8949 2.0089 1119.6 0.90
8 2.8953 3.0613 0.9735 0.3366 0.9606 57.1 0.52 2.1591 2.2597 57.1 0.80
9 3.2842 3.4362 0.9867 0.3522 1.155 31.1 0.42 2.3908 2.478 31.2 0.72
10 3.6375 3.7726 0.9926 0.3669 1.3396 105.3 0.35 2.5908 2.6652 104.9 0.65
11 3.9482 4.0643 0.9955 0.3798 1.5077 36.8 0.31 2.7612 2.8241 36.5 0.58
12 4.213 4.3101 0.997 0.3905 1.6538 52.8 0.29 2.9049 2.9578 52.5 0.54
13 4.434 4.5145 0.9978 0.3988 1.7766 43.6 0.27 3.0256 3.0698 43.3 0.50
14 4.6168 4.6831 0.9983 0.4052 1.8782 14 0.26 3.1262 3.163 13.9 0.47
15 4.767 4.8214 0.9986 0.41 1.9613 37.9 0.25 3.2099 3.2404 37.9 0.45
16 4.8896 4.9333 0.9989 0.4137 2.0289 30.9 0.24 3.2792 3.3044 30.5 0.44
17 4.9883 5.0235 0.999 0.4165 2.0831 1.5 0.24 3.3366 3.3574 1.4 0.42
18 5.0676 5.0958 0.9991 0.4187 2.1266 38.1 0.23 3.3837 3.4008 34.5 0.41
19 5.1312 5.1537 0.9992 0.4204 2.1612 6.9 0.23 3.4225 3.4364 5.7 0.40
20 5.182 5.2002 0.9993 0.4217 2.189 3.7 0.23 3.4542 3.4656 2.8 0.40
21 5.2888 5.2963 0.9994 0.4242 2.2466 27.8 0.22 3.518 3.5245 12.4 0.39

Relative fishing intensity by gear in 2006:

Trawl 23%, Hook&Line 15%, Set Net 1%, Recreational South 44%, Recreational North 17%

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