~~

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Written and drawn by Jay Hosler. (c) 2011 by Jay Hosler

 'glmme Sorne
sugar"
SíARRlN& ,,
@ WlLBUR &
ANTEDNA

Bf.CAUS 6 \F
"\'ou \iAD '{OU'D
l<NóW THAT PLANTS
DON'í Go OOZ.ING¡
'BuC.l¡(erFULLS o F
SUE:iAR ALL O'ltR
me flLACE.
 Wt-\AT 'PO
Yov t'15AN
"PASS?"

I'M NOTTPiLKINú A.BOUT

IlE\IE LO PIN Gi 'PlCfu RES ! I' n JUST LE'í
TALl<.1 N C:.\ ABovr TH'í= 'PROCE.S S ME €)(flAIN IT
TO '(O\J.
1HPrT Tl?f\1'5 A 11NY ?ACKET
OF Tl-lE Süt-.l'S E-NERfi'< IN
TI!t ltF€-SUSTA!NlN C::i.
MoLECULE oF {:¡LOCOS E
THAT MAKE.S OUR \JE.{<'{
E:)C.lSTENCE
?OSSlBLE.

REACtANTS

PRODUCTS

NOW AS 't'oll CAN SEE. 1 WATER) LIGHT AND

CARBON DlO)(IDE GO tNTO íHE FftOTOS'lNnl&llc.
PROCESS AND O~GEN, WATE.R. AND GLUCOSE CoMe ovr:
r'HofóS'fNTHES!S (5 conPoSED Of íWO SEPARATE S~íS OF C.HE:MlC.?\L REAC1lONS:
TI-!€ LIC:i.HT- DEP1=.NDENT RE.FICTlONS AND nH: LIGHT- /NDEPEN DENT R~ACT!ONS,

11-IE UGJ-IT-Df'PEND(;.NT REACTION5 USE n-lE Ll~HT­ 1NDfP€NDE"t­JT R6ACTioN5 Do

WAíeR AND ENERG"t FROM A PHorON OF NOT REQUIRE. Li<;HT DIRE.CTL'{. tN
LiGil-tT TO EUIL-b n-tE MOLf:CULES ATP 11-!ES E R EACllONS, CAR BON DIO~I D 6
AND NAOPl.J. O)('/C.EN rs RE.LEASE:D AS IS POLLED FROM n\E. AlR. AND
WASTE. ATP AND NAt>PH AR.E USED ATfACHED TO AN t::)(ISTING MOLf:ClJLE...
AS FUEL. FDR 1HE Ll~HT-INDE:P~NDENT 11-lEN THE .ATP AND Nt.WP\i ARc LJS(D TI>
REAcnoNS. 1ÜIZN Tl-JAT MOL,ECULES lNTO GLUCOSE.

WATl:R LlC:tHT
'' ..
. .'
.
'
.
~.: ···7 :. .. :

LIGWT­
DEPENDENT
REACTIONS
/:f@is ., ......
•. '
' . .·
..
i.7 -'. ~:.::.,· .... -··- . -,
.:
.r····· · · : :;
~-·:,::'.{:f;•
-: ~

! [i fi:.U\}:}1 Jtf 1; !i·}(C'.!~}[':11=;

1

i }: lE /\'/( T·' .....i] (~) iH.J ti:

.. ,
'.
.
:
,
~
.
:
.. ...
..... ·-. ···- ..
 So HOW DO
PLANTS CAíCl-t
A PHOiDN or
LIGHT?

,1···,,,
~'
0
----
~,1,ic;,.,
-
--
:

~ o
~
 \NSIDE n-tE CELLS ARE
E\/E:N SMALLER. FLulD-FILLED
BAEiS CALLED ORG.ANElLtS.

PHOíoS'/NTHESI S . OCCU RS
IN ORGANELLE:S CALLfD
CH LOROPLASTS. .

~BÉ PLANT CELL

NoLU, wHi DoN'T Yóu
KI\RA-1F oto P lrtl'::> Cl1LOIZo-
PlP&ff 1 N HAL'F.

nlE: Tl-l'(LAKOIC>S ARE.-

ARRAN~ED IN STAC'l.5 __,
CALLE.O GR.ANA.
( a $11"\iie s ~ acx Qf ti-,~\a\<,,,ds -
is Cf\L~O a '-RI\NUt-1)
 S'E\/ERAL PRolE:\NS AND
-....-...- GTHER MOLEClJLE5 RRE STuC.i< IN
-ruE THYLACOID MfMBRANE. UGif+T
!S CA PT\JRE.D B'{ CLUS1ERS O'F
CHLóROPH'/LL MOLECULE-5 IN n+E
MEMBRANE CALLED PHOTOS"/STE{Y)S.
~ tN PU:lNTS THE RE ARE TWo FoR THIS {::)(PLANAilON,
TYPES, CLEVER LY NAt"\EP WE1LL START WITI-1
PHoToS'fSTEM T ANP ?1-\oToS~STé.M Jl.
P ftOTo:S'(STEM 1f

SHOULDN'T
WE: SfART AT
?HOíoS'iSTE.M I
S!NCE, WfLL, IT
HAS nl-f NOMBER
ONE: IN rrs
NPIME?

AND ~OU'RE LEARNINC::i

ABOUT PHOTOS'/NTl-lESlS,
n~ W\N W\N!
íHE PHoToS'{Sí6M HAS ít.J o MAJoR REGWNS. IMA'=ilN E !"M A CHLoRoPH'fLL
nlE LA-R~E ?\l'(TENNA (011\PLE)( ~UR,ROtJNDS MOLEC.ULE AND '(O(.) ARE A SPE.CIAL
THE SMALL6R R.EAC.TfOt.l C€NTfR. E.NER~'{ . PAlR or MOLECUL&S ll'J
FRoM A PltOTDN OF LlEirtT IS ABSoRSf.D 1116- REA.CílON CENTE.R
8'( THE CHLOROPH'{LL IN THE ANTE-NNA CALLEO P680.
COMPL~)C AND FUNNELED TO ntE
RE-AC110N CE:Ní€R.

~!-/LO RoPHYlL]

Hl GH WHEN CHLORon-\'{ L~
fNER<H ABSORBS A PHOioN,íHE
ENE.R.~'{ E.Lt.VAÍÉS ONE:
OF 11-{E, G\loRoPH'/LL'S
NOW, IMAGII\IE ELECfRoN~ To A HIGHER
OUR \-tÉADS ENERGi'I STA-TE:,
RE.PRE:SE~T
AN. EL.ECTRON
ON íHE
MOLECVL.6.

LOW
E_('!_E_R_s:i:x___
H!~H H\GH
ENER~Y _ ENER.C.'( _

... AND \í
.. vNnL
!T \5
PASSES THE E:\/ ENíV A Ll '{
EN.ERGi'{ To A PA-SSE\:> TO
NElf:\1-tBORlN (;¡ P(é,80
CHLOROP\i'/LL, IN 11-\e.
E~!TINC:¡ \íS . Rc:ACTION
E.LECT?..ON. CENTE-R.

Pf>SO } P~So ·

..
)
- - - - - .. - - - - - - -- '-.) (;, - - - - -- - - - - - - - - - - - -- - -- ---- --

)"' '<oP/ (t_

i""' ('
 íHE:RE 'S ~ REGloN
TO RSPLACE ntE
or PHOíOS"ISTéM 1I
CA-LLeD n+E- O~'f&EN
ELECTRONLOST BY
P6'&0 lN PHOTOS'ISTfJ1 JI,
E:\fOL\llN& CoMPLE)( «ie.c)
WE W lLL NE.ED TO TAKE. í1-IAT SPLITS WATER
ONE. f'ROM ANOn-t~R I NTO 11\REE 'PA RTS ;
MOLEC.UL.E. TN 11HS (..\\/ DROGEN, 0"1-'iGf:N &_O('JE
CASE, 11-\AT OTHER Of Tl-lE ELECTRoNS
MOLECULE:; is -nlflT KN rrs THE M
WATER. T?>Gf:":THE~.
 'lou SA!D LJ~Hf-11EA06D AND
WPRE TALKlNG¡ Ai30Uí
PHoroS'fNlHES!S AND 1HA1 US€5
LlG,l-tT •.. AND .•• Uh-- 'foUr nea<I
... (S ... an e\ectron wn,ch has ...
... a1111ost no rnass ...
,- So Lts ...

PS '{OU RECALL> THE ELECn'ON

fl<OM P"80 WAS CARRlE:D A\JJFt'{
11-tE LOMEN
®
By PLASíoQU!NONE. PQ TAKéS AlREAD'í H-A5 f\
11-tE ENERG¡r2.ED E LECTI<,ON TO BUNc..H- OF b';(Tl<.A
11-iE C'/TOC.ttRO ME CoM P L&)<. 1-l'f DROGSN FROl-'1
WHlCH OSE:S 1HE ELECTRON'.S 11-tS WATBR- VJE
5")(.TRA E.NERGt'< W -PUM? S\>L-l T.
MORE. HYDRO(;E.N lNTD
THE LUMEN.
 o,'
1
"-...

TI-tERE ARE LO-r-5 oF HXD~OG.&N !ONS IT TAK8S nte HARD WOR.K. OF íH€
ll'{SlDE. 11-lE LLIME:N, BVT NOT '\/ER'f MAN'1' C'IToCl-lROME CCit1PlE)<' TO MA-INTAIN n+E
IN 1HE: SURROVNDING, STRoMA. TH-E. CoNCE~AllON G'RA-Dl&Nr. IN NA-TUR~.11-llN~
DIFFERENCE. lN CONCf:NTR..ATION ACROSS Ul<E To SPRGA-D n\E't'\SEL\/ES OUT SVENL'{.
n-t~ nl"ILA Ko10 MEMBRAN0 IS CALLE:D GNSN A CHANCE., THE lf'IPROG.EN lONS
A CONCENTRATION GRAt>IENT. C.RAMM.E-D JNTO Tl+E LVM5N WO UL D
-~- +--.;;:::-- . CROSS THE l1"\'ILA !<.DI D
MEHSRA:NS A-ND
SPRtAV oor IN TI)
TltS STFD MA.

ALL Clf THt YEAH, \JfJ..L, 1cE1WEfN Tl-\E

Po51T1VE:-L Y CHAR~ED CO"!Cf NTRATlON GR,A'Dl'cN T
HYDR.OqEN \ONS ARE AND AL.l of 11-\~lR i>OSl T"l\/E.
CHARG.E-5, Tl-tE ~iDR.OCeiE-N !ON'S
PACK'eD IN TIGliTL'( RE.AW{ WANT 10 L-€AVE
AND TltE-lR PoSlTIVE 11-tf; LUMEN AND 5PREA-D
CHAR,GiES REPEL O \JT l NTD ntE <;;"TROM A.
f.AC.H an, ER.
1-lowEVf:R, TI-,(: ONLY
í)'VE. Of DOOR ACROSS
ntE 11-l'llfl KOID MEMBRANE.
11-\E.'{ CAN USE 15 R
PRoTEIN CAl.LED
AíP S'INTHASÉ.
 OH> 'iOUR HfAD
(AKA, rue ELECTRON fR,OM
PHOíCl5'/Sí~M rr) REnJRNED ío
\íS LO//JER E:NERG't' SíATE .
A-Ftt-R \T E:N€R~lZ.8D
11-tE C'{Tt>CHROME
(oMPLE)(.

NOW WE NEE\) To TuRN OIJR LET'S Do A urn,e ROLE

AíTtN1lON To Pt­loToS'f5T€M 1. PL.A'/IN~. T'Ll BE: A
LIKE.. ?51l,1)ttc:rroS'ISTt.M I CHLOROPHLL MOLECULE IN
ASsoRBS A 'PHOíoN AND THE ANTE.NNA Co{'i)Pl.E;,< OF' ?SI
VSES 11-lE EN 6RC:i'< To AND vouCAN BE. P100, A
DONATE AN ELECfRON SP€C.tAL PA\R OF MOLECULES
TO ANOT\-lE.R IN n-ti; REAC1\óN
MOL'E.ClJL.E. CE:N~R-
 THE Cf-lLORoP~'iLL. WHEN P7oo A85oRB5 IN FACT, IT GETS So
MO LE:CULE.S IN 11-lE Tl-tE é:NERGi"(- E)(C\TED rr POPS OFF ANP
At{TENNA (OMPLE)( OF RlDE=S AWA'í ON A MOLECULE
PSI ABSORB A PH010N'S CALLED FERREDO)UN.
ENE.Re;'< AND PASS
!T AROUND LJNTI L
THG'f PASS \T
To P700 lN T4E
RE.AC.TION
CE.NTER.

3 ..... - ..
( Cf.lLOR_oPHYLl}

fe;RREDO't(JN TA KE5 THE

ELE:.CTRON FROM P70o To A
M OLECULE CALLED NflDP" REDUCTASE..
NADPt- REDVCTAS E USES TI! E
:~LELTRON'5 ENER(i'( To
MAKE A MOu:;cvu:
OF NADPH.

Of CoURSE,
NOW P~Oío5'f5ITM I I DIDN'T CARH
NE.tDS To REPUlCE ITS 11-IAT DID '(ov SAY
LOST ELt:C.TRDN. C A-N "FROM SPL11TlNGi
'iou f¡U~SS WHERE A V\/Aí~R MOLfCULE?''
lBAT l<,EPLACEMENT
8E.CAV5 E
COMES FROM? 11-Htf is
NOT ntE
ANSWER.
 WHEN T\f E PooR POOPE:'.D tLECTRON FR.OM
F'HOTcS)'STEM 1t L~A'JfS rue C'lTOCHROME
(OMPLBX, lT H 11t.H es A RID E ON A
MOLECULE. CALLED PLASTOC'{ANIN (Pe.).
Pe íAKE-5 ll-\E HECTRON lb PSI ...

..• WHE..RE IT Rlfi/tT. BUT ntG Llfiltt

GRE:i,T. /NDePENDEN T Rrnc.110N S
REPLACE5 11-IE' SOiNOl>.1
¡; LECTf<ON 11-\AT WE:. HA'IE TAKf PLACe JN TI-}E 'STRo/l111\,
So WE' 5!,0UL P G~r cor
PHDToS'f S TE;. M r n-tE ATP OF 1--tÉRE..
LOSí. AND NAUPH
fOR ntE
UC:i,H-T-
;I N Df PE:N Df.NT
'REA-C.nON S,
Rl~ftT?

OH, WEL.l>
A LL OF TI-! ES E.
H'iOROG.EN I ONS
MAKE. Tl-lE LUnEN
Of TI-tf nJ'{LAko!D
\>RE;"ft'{ acro 1 C.
 Ros1sco coMB1NE-S "BEU\VSE.
CO;. . wm-1 A FIVE SU~AR is A RtNGi
CARSóN MoLEüJLE OF CAR\30N. !F 11-lE
CALL(:;D RuBP IN A Lll:iHT-INOEPEDfNT l<~AalON~
PRoCESS KNoWN t\-S ARE ~ONNA MAl-(E SVGA-R, .
~CARBON F"n<A110N. '1 11-!E'{' \JE ~OT 1D G~T n\f:
CARBON BVtLDINGi 'BLOC.KS

----------
FR.OM 5oM6WHER€·
 So. WAlT. IF
Rus1sco PVLLS
C02 FR,01'1 ntE
A1R. Wl-lERE
noss IT ~ET
RuBP?

-the
Calv¡n
Cyde

ONE OUT
OF E.V~R'i St-X
~3 p Mo LEC(}LES
MANUFACTURE. D
LEA\f(S THE C"{CLE
AND G0125 oN
To MA 1<.E.
SUGAR.
 .-
t'LASToC.YANIN
IS TrfE 13RTDGE: CoNNE'.CTrNG
t TH( 1:HoTOSY5TEM.S
 T€C.Ht,¡1CALL'I
G3P lSTI--tE END
OF 1'HcríOSYNT4E:S\S
ANP Tl-!Aí1s ALL
I SAltl Wf1D
TALK ABOVT:
I 'JYI
NOT Rt:AU~
FAMILtAR
WIT1-t nte
°BlOOtEMlCA'L
PATH WA'fS
11-lAT (ON\lCRT
GioP INTb
GLll(QSE:
 GEE-Wl·H'Z-,
WIL.6UR. [F You
REALL'/ WANTED
SoMETHING
5WE.GcT you
COU1.-D I-IAVE
CONJUR[D rr
AN'/11 M t-·

+ ~J>P­•,

I JuST Wf.NT
íO A LOT OF íROVBL.E-
To 8;<Pt.AIN SOMfTHlNG
AM~21N&.