14 Ecosystem
14 Ecosystem
Forest
14.1 ECOSYSTEM-STRUCTURE
AND FUNCTION
Each ecosystem is composed of both structural
and functional components, and operates as
an open system that receives input in the form
of solar energy.
The flow of energy within an ecosystem is
unidirectional, moving from producers to
consumers i.e, towards the higher trophic level.
The dissipation and loss occur as heat to the
environment.
COMPONENTS OF ECOSYSTEM
Cycling of matter occurs between the biotic and
abiotic components of the ecosystem.
1. BIOTIC COMPONENTS
Biotic components make up all living members
of an ecosystem and are connected through
food and various other relationships.
(A) Producers/Autotrophs
(B) Consumers/Heterotrophs
(C) Decomposers
2. ABIOTIC COMPONENTS
Abiotic component of an ecosystem consists of
non-living substances and factors. The
important ones are as follows: Temperature;
light; wind; humidity; precipitation and water.
ECOSYSTEM STRUCTURE
Interaction of biotic and abiotic components
results in a physical structure that is
characteristic for each type of ecosystem.
SPECIES COMPOSITION:
Ecosystems are studied by identifying and
counting the different plant and animal species
that exist within them.
The species composition varies between
ecosystems and is determined by geographical,
topographical, and climatic factors.
The highest diversity of species can be found in
tropical rainforests and coral reefs, while the
lowest is in deserts and the Arctic.
STRATIFICATION:
Stratification refers to the formation of vertical
layers of vegetation, such as in tropical
rainforests where there are 5-7 strata with tall
trees, canopy trees, and understory trees at the
top, a shrub layer below the tree layer, and a
bottom layer of herbs and grass at the ground
level.
Stratification
TROPHIC STRUCTURE:
Each ecosystem has specific food chains and
food webs, e.g., grazing food chain in grassland.
Standing Crop:
Standing State:
PRIMARY PRODUCTIVITY
The amount of energy accumulation in green
plants as biomass or organic matter per unit
area over a time period through the process of
photosynthesis is known as primary
productivity.
It is of two sub-types.
(i) Gross Primary Productivity (GPP):
SECONDARY PRODUCTIVITY
Secondary productivity refers to the rate of
formation organic matter by consumer
population (NEET 2013).
It is influenced by the loss of energy-rich organic
matter during transfer from one trophic level to
the next, as well as consumption through
respiration and predation.
Respiration losses can reach up to 20% for
autotrophs, 30% for herbivores, and 60% for
carnivores, causing a decrease in net
productivity with each increasing trophic level.
PROCESSES INVOLVED IN
DECOMPOSITION
1. Detritus Fragmentation: Detrivores (e.g.,
termites, carrion beetles, earthworms) consume
larger pieces, leaving smaller fragments behind
(NEET 2013, 2022, 2023). This also occurs when
detritus passes through the digestive tracts of
animals. This process increases the surface area
of the detritus particles.
2. Leaching: Water that percolates through the
soil carries soluble substances like sugars and
inorganic nutrients to deeper layers of soil or
ground water (NEET 2023) and get precipitated
as unavailable salt.
3. Catabolism: Decomposers like bacteria and
fungi excrete digestive enzymes on the detritus.
This changes complex organic substances into
simple and soluble organic compounds and
inorganic substances. Some of the broken-down
food is taken up by decomposers and
immobilized.
4. Humification: Humification causes the
accumulation of a dark-colored, amorphous
substance called humus. Humus is rich in lignin
and cellulose and is highly resistant to microbial
action, which leads to slow decomposition. It is a
reservoir of nutrients and helps maintain soil
moisture and aeration. Humus is colloidal in
nature.
5. Mineralization: Some microbes further degrade
humus (NEET 2023), which leads to the release
of inorganic nutrients through the process of
mineralization. It is a slow process because the
nutrients are trapped in humus and immobilized
in decomposers.
Fig. 14.3.1 : Decomposition process
FOOD CHAIN
A food chain is a sequence of nutrient and
energy transfer in the form of food from one
organism to another within an ecosystem.
This involves repeated eating, with each group
being eaten by another and passing on the food
energy.
Typically, a food chain is straight and consists of
populations related by eating and being eaten,
with 3-6 trophic levels.
Humans, however, can occupy more than one
trophic level. As the trophic level rises, there is a
progressive reduction in available biomass,
energy, and number of individuals.
Most of the energy at each trophic level is lost as
heat. Producers and decomposers sustain food
chains.
Trophic level refers to the specific place in a
food chain where an organism obtains its food.
The number of trophic levels is equal to the
number of steps in the food chain.
The amount of living biomass or number of
organisms in a unit area of an ecosystem or
trophic level is called standing crop, which is
usually measured in dry weight to avoid
seasonal moisture differences.
(
it follows the 10% law (only 10% of energy is
transferred to each trophic level from the lower
trophic level) (AIPMT 2014).
1. PYRAMID OF NUMBERS
An ecological pyramid displays the number of
individuals per unit area in each level of the
food chain, with producers at the bottom and
top carnivores at the top.
In most cases, the pyramid of number is upright,
with fewer individuals in each higher trophic
level.
The largest number of individuals is found at the
producer level, and successively fewer
individuals are found in herbivores, primary
carnivores, and finally, top carnivores, which
have the smallest number.
Fig. 14.5.2 : Pyramid of number in grassland
2. PYRAMID OF BIOMASS:
The amount of living organic matter is referred
to as biomass and can be measured as fresh or
dry weight.
The pyramid of biomass is a visual
representation of the biomass present at
different trophic levels per unit area, with
producers at the base and top carnivores at the
tip.
3. PYRAMID OF ENERGY
LIMITATIONS OF ECOLOGICAL
PYRAMIDS
The concept of ecological pyramids is based on
the idea of simple food chains, which are not
found in reality as complex food webs exist in
nature.
Ecological pyramids cannot accommodate
cases where a single species functions at
multiple trophic levels.
These pyramids do not take into consideration
the significance of detrivores and decomposers
in the ecosystem.
14.6 ECOLOGICAL
SUCCESSION
Each community undergoes constant change in
response to changing environmental conditions,
with this change occurring in an orderly and
sequential manner that parallels changes in the
physical environment.
Ecological succession refers to the gradual and
predictable changes in the species composition
of an area, during which some species become
more numerous while others decline or
disappear.
These changes occur in a series of transitional
communities known as seral stages or seral
communities, with the initial group of organisms
colonizing bare land known as pioneers or
primary colonizers and the final group of
organisms in a stable, self-sustaining biotic
community at the end of succession referred to
as a climax community. The entire sequence of
community that successively change in a given
area are called sere(s).
The climax community is in equilibrium with its
physical environment and has maximum
diversity and niche specialization. As the
diversity and number of species and organisms
increase, so too does the total biomass.
The communities that exist today are the result
of millions of years of succession, with evolution
and succession occurring in parallel during this
time.
TYPES OF ECOLOGICAL
SUCCESSION
Ecological or biotic succession is divided into
two types: primary and secondary succession.
Primary Succession: This type of succession
occurs in an area that was previously barren or
lacked life, such as newly exposed sea floor,
igneous rocks, sand dunes, new cooled lava
sediments, or newly created pond or reservoir.
The environment is hostile to the first life or
pioneer community and the process takes a
long time, often thousands of years, as soil,
which is necessary for establishing a biotic
community, takes several hundred to several
thousand years to develop, depending on the
substrate and climate.
Fig. 14.6.1 : Primary Succession
IMPORTANCE OF ECOLOGICAL
SUCCESSION
The sequence of biotic succession is usually set
and ecologists can easily determine the seral
stage of a biotic community in an area.
Understanding biotic succession allows us to
maintain certain stages of development, such
as a pasture composed of grasses and herbs,
by interfering with the progression of succession
through activities like grazing and fire.
14.7 NUTRIENTS CYCLING
Organisms require a constant supply of
nutrients in order to grow, reproduce, and carry
out various bodily functions.
The amount of nutrients, such as carbon,
nitrogen, phosphorus, and calcium, present in
the soil at any given time is referred to as the
standing state, which varies in different types of
ecosystems as well as seasonally.
Nutrients are not lost from ecosystems but
rather recycled indefinitely through a process
called nutrient cycling or biogeochemical
cycles.
Nutrient cycles are classified into two types:
Gaseous and sedimentary.
PHOSPHORUS CYCLE
Phosphorus is a vital component in biological
systems, participating in metabolic reactions
that release energy from food and aiding in its
utilization in various functions of the body.
It plays a crucial role in forming nucleic acids,
bio-membranes as phospholipids, cellular
energy transfer as ATP, and structural elements
like shells, bones, and teeth.
Phosphorus primarily exists in the form of
phosphate and has a reservoir pool in
phosphate rocks and a cycling pool in soil in
terrestrial ecosystems, and in water in aquatic
ecosystems (NEET 2013).
The cycling pool is replenished through the
( )
weathering of rocks (NEET 2022), and phosphate
is typically found in soil in combination with
calcium, iron, and aluminum.