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Monograph 1

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Atashi Mandal
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Phytokeys 37:25-101 (2014) A peer-reviewed open-access journal

MONOGRAPH
Doi: 10.3897/phytokeys.37.6499
Launched to accelerate biodiversity research
www.phytokeys.com

Revisions and Key to the Vernonieae


(Compositae) of Thailand

Sukhonthip Bunwang 1, Pranom Chantaranothai2, Sterling C. Keeley3

1 Maejo University Phrae Campus, Mae SAi, Rong Kwang, Phrae 54141, Thailand 2 Applied Taxonomis
Research Center, DFEpartment of Biology, Faculty of Sciene, Khon Kaen University, Khon Kaen 40002,
Thailand 3 Department of Botany, University of Hawaii, Honolulu, HI 96816 USA

Corresponding author: Sukhonthip Bunwang ([email protected])

Academic editor: A. Sennikov / Received 25 Octbor 2013 / Accepted 4 April 2014 / Published 13 May 2014

Citation: Bunwang S, Chantaranothai P, Keeley SC (2014) Revisions and key to the Veronieae (Compositae) of
Thailand. Phytokeys 37:25-101.doi: 10.3897/phytokeys.37.6499

Abstract
Seventeen genera and 48 species, in five subtribes, are recognized in Thailand. These include 15 endemic taxa,
half of which are in the largest genus Acilepis, with others in the genera Camchaya, Koyamasia, and Okia. A
new monotypic genus, Pulicarioides, is established with P. annamica, the new name for the species formerly
known as Vernonia pulicarioides. New combinations are also made for Acilepis kerrii, Cyanthillium montanum,
Koyamasis curtisii and Okia pseudobirmanica. Forty-six characters of taxa were identified. Keys to achene and
pollen morphology were analysed using UPGMA. Five clusters of taxa were identified. Keys to genera, species
and varities, descriptions, vernacular names, ecological data nad illustrations are provided.

Keywords
Acilepis, Asteraceae, Camchaya, Cichorioideae, Cyanthillium, Decaneuropsis, Elephantopus, Ethulia,
Gymnanthemum, Iodocephalopsis, Koyamasis, Kurziella, Monosis, Okia, Pseudelephantopus, Pulicarioidea,
south-east Asia, Strobocalyx, Struchium, Tarlomounia

Overview of the Vernonieae and taxonomic history of the tribe in Thailand

The Vernonieae have been shown to be unequivocally monophyletic and are well represented in
both the Old and New Worlds, with centers of diversity in east Africa and Brazil (Keeley et al. 2007,
Keeley and Robinson 2009). Recent revisions by Robinson (1999a, b, 2007) based on morphology,
secondary chemistry and palynological studies,

Copyright Sukhonthip Bunwong et al. This is an open article distributed under the terms of the Creative Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and
reproduction in any medium, provide the original author

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