Ecology Assignment
Ecology Assignment
Submitted To Submitted By
Md. Rakibul Hassan Asadul Islam
Assistant Professor & Head of the
Roll: 22103403
Department,
Reg: 10894
Department of Environmental Science and
Session: 2021-22
Engineering
2 year 1st Semester
nd
Jatiya Kabi Kazi Nazrul Islam University
Department of Environmental Science and
Engineering
Jatiya Kabi Kazi Nazrul Islam University
List of Tables
Table 5.1-I: Energy Distribution of Solar Radiation on the Earth ............................................. 3
Table 5.2-I:Forms of Annual Primary Productivity for Several Ecosystems (kcal/m2) ............. 5
1. Introduction
Energy flow through ecosystems is a
fundamental process that maintains life on
Earth. It begins with solar energy, which is
captured by primary producers, and is then
transferred through various trophic levels in a
series of steps known as food chains or food
webs. In ecosystems, energy is neither created
nor destroyed but transferred from one
organism to another, following the first law of
thermodynamics. From an engineering and
environmental science perspective,
understanding the flow of energy is critical for
designing sustainable ecosystems, assessing
environmental impacts, and applying these
principles in real-world solutions, such as
Figure 1-A: Sundarban, The largest Ecosystem of
biomass production.
Bangladesh
In Bangladesh we have the largest mangrove
forest in the world, which is our largest and enriched ecosystem. To protect it, we have to
learn about the energy flow and the ecosystem mechanisms.
• To explore how energy is transferred across trophic levels in food chains and webs.
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3. Theoretical Framework
3.1. Laws of Thermodynamics in Ecosystems
The flow of energy in ecosystems is governed by the laws of thermodynamics, particularly
the first and second laws. The First Law of Thermodynamics, also known as the law of
energy conservation, states that energy cannot be created or destroyed, only transformed. In
ecosystems, solar energy is transformed into chemical energy through the process of
photosynthesis. This energy is subsequently transferred to other organisms in the form of
food, but no new energy is created at each step.
The Second Law of Thermodynamics explains that in every energy transfer, some energy is
lost as heat. This law introduces the concept of entropy, which suggests that energy quality
decreases as it moves through trophic levels. The energy that powers the activities of higher-
level consumers (such as carnivores) is therefore much reduced compared to the energy
initially captured by producers. Understanding these laws is crucial for ecological
engineering because they help explain the inefficiency of energy transfer and the limitations
this imposes on ecosystem structure.
• The Y-shaped model provides a more accurate depiction by accounting for two
parallel channels of energy flow: one that goes through herbivores and their predators,
and another through decomposers. This model better represents the interconnected
nature of ecosystems and provides a more realistic framework for studying energy
dynamics.
These models help environmental engineers and ecologists calculate energy efficiency and
understand how energy distribution affects ecosystem sustainability.
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5. Flow of Energy Through the Ecosystem
5.1. Energy Sources
The primary source of energy for most ecosystems is the sun, which drives the process of
photosynthesis. Through this process, plants, algae, and some bacteria convert solar energy
into chemical energy in the form of glucose. This chemical energy is stored in the bonds of
organic molecules and becomes available for consumption by other organisms in the
ecosystem. Geothermal energy derives from radio-active decay in the Earth and contributes
about 0.5% of the solar input. Tidal friction extracts energy from the kinetic energy of the
Earth–sun–moon system and is about 0.0017% of solar. A portion of the wind’s energy comes
from the kinetic energy of Earth’s rotation. Oxidation of reduced inorganic minerals
transported to the biosphere from deep in Earth’s crust by hot springs and deep ocean vents
provide energy for unique ecosystems adjacent to them. (However, this source is not
completely independent of solar input, as it relies on oxygen produced mostly in the
photosphere by solar energy.) These are the ultimate sources of energy for the biosphere.
However, sensible heat and mechanical forms of energy cannot be used to form
carbohydrates from CO2. Organisms require high-quality energy in the form of photons or
chemical bond energy in order to drive the reactions that produce biomass from inorganic
precursors.
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5.2. Primary Productivity
Most of the solar energy that reaches the Earth's surface is either turned into heat or used for
processes like evaporation, wind, and waves. Only about 0.8% of this energy is captured by
plants and other organisms through photosynthesis. This process provides almost all the
energy needed for life on Earth. Organisms that perform photosynthesis are called producers
and they fix carbon from CO2 to create organic matter. The speed at which they do this is
called primary productivity, and it can be measured in terms of energy (kcal/m² per day) or
biomass (g/m² per year). Biomass refers to the dry weight of these organisms, and
carbohydrates provide about 4.1 kcal/g.
However, not all energy-fixing processes are considered primary production. For instance,
chemoautotrophs, which live in places like ocean vents or marshes, get their energy from
chemicals like hydrogen sulphide. These chemicals often come from other organisms or non-
living sources like volcanoes.
Primary productivity has two parts: Gross Primary Productivity (GPP) is the total amount of
carbon fixed by producers, while Net Primary Productivity (NPP) is what's left after the
producers use some of that energy for their own respiration. In other words, NPP is the
energy that gets passed on to other parts of the ecosystem.
GPP can be estimated by measuring NPP and respiration. In some environments, like oceans,
GPP is closely tied to the amount of chlorophyll, the pigment in plants that captures light for
photosynthesis. Chlorophyll can be measured, and then a factor called the assimilation ratio
(the rate of carbon fixation per gram of chlorophyll) is used to estimate GPP. This ratio varies
by ecosystem, but in the ocean, it's roughly 3.7 grams of carbon fixed per hour per gram of
chlorophyll. In forests, this ratio can be lower because plants have more chlorophyll to
capture light. In sparsely vegetated areas, there is less chlorophyll, but it fixes carbon at a
faster rate per gram.
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Table 5.2-I:Forms of Annual Primary Productivity for Several Ecosystems (kcal/m2)
It is important to understand the relationship between the gross and net productivity, which
are rates, and the biomass, or standing crop. Keep in mind that the amount of biomass present
is not a measure of the gross or net productivity. As an analogy, consider a tank with water
being pumped in, and with some flowing out through a hole in the bottom. The inflow is like
the gross primary productivity, and the outflow is like respiration. If more is entering the tank
than is draining out the hole, the difference accumulates. The rate of accumulation is like the
net primary productivity. If the amount entering and leaving are equal, the amount in the tank
stays the same and the system is at steady state. This would be equivalent to having zero net
productivity and could occur with the tank full or almost empty. Consider a mature tropical
rain forest (Table 1). The gross primary productivity is of such a forest is about 45,000
kcal/m2. yr. However, most of that is taken up by respiration to maintain the large biomass.
Thus, the net primary productivity is essentially zero. On the other hand, an alfalfa field is
adding considerably to its biomass during the growing season. Its gross primary productivity
is 24,400 kcal/m2. yr. Respiration accounts for 9200 kcal/m2. yr, leaving a net primary
productivity of 15,200 kcal/ m2. yr. However, the actual yield that farmers can obtain from
the field will be lower because other heterotrophs (such as field mice) consume part of the
crop. This leads to a third type of productivity, net community productivity (NCP), which is
the gross primary productivity minus plant (autotrophic) respiration and minus the amount
consumed by other organisms in the ecosystem (heterotrophic respiration). In the case of the
alfalfa field, 800 kcal/m2. yr is consumed by heterotrophic respiration. This yields a net
community productivity of 14,400 kcal/m2. yr, for an efficiency of 59%. Primary productivity
(GPP) can be limited by numerous factors, such as the amount of sunlight, availability of
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moisture and nutrients, soil properties, and so on. On land, moisture is the most serious
limiting factor. In the open ocean, productivity is limited by nutrient availability and the
ability of sunlight to penetrate the water. Ideal conditions can result in productivities that are
as much as double the ‘‘typical’’ values; 50,000 kcal/m2. yr is thought to be the practical
upper limit for any natural ecosystem. Changes in primary productivity, or comparisons to
similar ecosystems, can be used as indicators of the health of an ecosystem. Worldwide, the
total GPP is estimated to be about 10 18 kcal/yr. The primary productivity of the world’s
agriculture has only been keeping pace with increases in human population. Humans greatly
increase the productivity of cultivated land by energy subsidies. This energy originates
mostly from fossil fuels and takes the form of synthetic nitrogen fertilizer, pesticides, and
mechanized agriculture (tilling, harvesting, etc.). Productivity of crops has also been
increased by breeding new varieties. The greatly increased food production has resulted in an
increase in the carrying capacity of the Earth for humans, and has been called the green
revolution. However, it must be noted that the new crops are more dependent on energy
subsidies than are traditional cultivars. Although the improved productivity has been
important, underdeveloped countries have increased their food production as much by
increasing the amount of land under cultivation. This results in destruction of forests and
resulting loss in habitat for other species. Humans have other uses for primary productivity
besides as food. Cotton and flax Fibers are used for clothing and wood for structures, paper,
and fuel. Overharvesting of wood in poor regions of the world has led to deforestation and
erosion.
• Producers (Autotrophs): These include plants, algae, and some bacteria that convert
solar or chemical energy into organic matter through photosynthesis or
chemosynthesis.
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• Decomposers (Detritivores): Decomposers, such as fungi, bacteria, and certain
insects, break down dead organic matter and recycle nutrients back into the
ecosystem. Energy that was stored in dead plants and animals is partially released as
heat and partially made available for other organisms to utilize.
Figure 5-A: Flow of energy and cycling of materials through a typical food chain
As energy moves from one trophic level to the next, a significant portion of it is lost as heat
through metabolic processes like respiration. On average, only about 10% of the energy at
one trophic level is passed on to the next. This is known as the 10% Rule and explains why
ecosystems cannot support a large number of high-level predators: energy availability
diminishes with each step up the trophic hierarchy.
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5.4. Food Chains
Energy flow through ecosystems can be simplified into two main types of food chains:
• Grazing Food Chain: This begins with producers and involves herbivores and
carnivores. In terrestrial ecosystems, for example, energy flows from plants to
herbivores like deer and then to predators like wolves.
• Detritus Food Chain: This chain is based on the breakdown of dead organic matter
by decomposers. Energy from detritus flows to detritivores like earthworms and
fungi, and then to secondary consumers. This type of chain is common in ecosystems
with significant amounts of organic material, such as forests and wetlands.
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Food webs illustrate that many organisms feed at more than one trophic level. For instance,
omnivores such as bears may feed on both plants (producers) and animals (consumers), and
decomposers can interact with multiple levels of the food web. This complexity enhances
ecosystem resilience by allowing alternative pathways for energy flow if one part of the web
is disrupted.
The total energy a trophic level receives is called gross energy intake. Some of this energy is
not used and gets eliminated as feces, which goes into the detritus food chain. The remaining
energy is absorbed, but part of it is excreted through urine. A lot of the absorbed energy is
used for breathing and staying active. The rest is used for growth and reproduction, which is
called the net productivity of that trophic level. The production efficiency (also called transfer
efficiency) of a species or of a trophic level as a whole can be defined as:
𝑛𝑒𝑡 𝑝𝑟𝑜𝑑𝑢𝑐𝑡𝑖𝑣𝑖𝑡𝑦
𝑃𝑟𝑜𝑑𝑢𝑐𝑡𝑖𝑜𝑛 𝑒𝑓𝑓𝑖𝑐𝑖𝑒𝑛𝑐𝑦 = … … … … … (1)
𝑎𝑠𝑠𝑖𝑚𝑖𝑙𝑎𝑡𝑖𝑜𝑛
Birds 1.3%
Warm-blooded animals seem to pay a penalty in efficiency. The rate of respiration in animals
can be related to body mass empirically by a log-log relationship. For example, for
herbivorous mammals:
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Where R is the respiration rate in kJ/day per individual and M is the body mass in grams (1
kJ= 0.239 kcal). Cold-blooded animals may use more than an order of magnitude less energy,
but the value will be strongly dependent on ambient temperature. Although they vary widely,
a rule of thumb is that the production efficiency of trophic levels is typically about 10%.
Thus, if the gross primary productivity is 60 kcal/m2.day, the net primary productivity would
be estimated at 6 kcal/m2.day, and the herbivores would have a net productivity of about 0.6
kcal/m2.day. Carnivores typically have a 20% efficiency, so net tertiary productivity would be
estimated at 0.12 kcal/m2.day.
At each trophic level, organisms must perform essential life functions such as movement,
growth, reproduction, and maintaining homeostasis. To carry out these functions, they rely on
respiration—a biological process in which food is broken down to release energy. During
respiration, a large portion of the energy consumed by an organism is used to power its
cellular activities. This energy is not stored within the organism's body but is instead
expended to maintain basic functions like locomotion, muscle contraction, cell repair, and
thermoregulation in warm-blooded animals.
For example, in mammals and birds, which are endothermic, maintaining body temperature
demands significant energy, leading to higher energy consumption compared to cold-blooded
animals. The energy that is used in respiration is effectively "lost" from the perspective of the
next trophic level because it cannot be passed on in the form of biomass. Only a small
fraction of the energy consumed is converted into new tissue (biomass) that can be used by
the next organism in the food chain. As a result, this process contributes to the significant
reduction in the amount of energy available to higher trophic levels.
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6.2. Energy Loss through Heat
According to the Second Law of Thermodynamics, energy transformations are never 100%
efficient, and a portion of energy is inevitably lost as heat in any biological process. This is
particularly evident in metabolic activities, where organisms convert the chemical energy
stored in food into energy that fuels life processes. During these transformations, some
energy is dissipated as heat, which is released into the environment.
This heat energy cannot be captured or used by other organisms within the ecosystem,
making it an unrecoverable loss. Consequently, as organisms at higher trophic levels
metabolize their food, less energy remains available for growth and reproduction, further
limiting the energy that can be passed on. Heat loss is one of the primary reasons why energy
flow in ecosystems is considered unidirectional—once energy has been dissipated as heat, it
exits the ecosystem's energy cycle entirely.
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When herbivores consume plants, only about 10% of the energy stored in the plant biomass is
transferred to their bodies in the form of usable energy. The remaining 90% is lost due to
respiration, metabolic heat production, and incomplete digestion. The same pattern continues
for secondary and tertiary consumers—carnivores that consume herbivores or other
carnivores retain only around 10% of the energy from their prey. As a result, the efficiency of
energy transfer diminishes with each successive trophic level.
This inefficiency explains why ecosystems typically have far more producers than
consumers, and why the numbers of top-level predators, such as lions or eagles, are relatively
small. The structure of the ecosystem reflects this energy imbalance, with large populations
of plants supporting smaller populations of herbivores, which in turn support even smaller
populations of predators. This decrease in energy availability limits the number of trophic
levels in an ecosystem, as there is simply not enough energy to sustain large populations at
the top of the food chain.
7. Ecological Pyramids
Ecological pyramids represent the flow of energy, biomass, or organism numbers within an
ecosystem. These pyramids help to visually illustrate the diminishing energy as it moves
through trophic levels.
• Pyramid of Biomass: This pyramid illustrates the total biomass present at each
trophic level. In many terrestrial ecosystems, the biomass is largest at the producer
level, while in aquatic systems, the biomass pyramid can sometimes be inverted due
to the high turnover rate of primary producers such as phytoplankton. [Biomass is
defined as the total dry weight of the total amount of living materials presented at a
trophic level]
• Pyramid of Energy: The energy pyramid displays the amount of energy present at
each trophic level. As you move up the pyramid from producers to top-level
carnivores, the available energy decreases significantly. This pyramid always has a
broad base, representing the energy captured by producers. it overcomes the difficulty
encountered in pyramids of number and biomass; i.e. inverted pyramid never occurred
here. It shows the total amount of energy utilized by the organisms in different trophic
level in a square metre, over a given period of time, i.e. it shows the number of new
tissues of organisms produced in a unit time
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Figure 8-A: Grassland Food Web
A study of a grassland ecosystem in the Midwest U.S. demonstrated that energy transfer
between trophic levels was highly dependent on plant productivity and climate conditions
(Fath & Patten, 1999). Changes in precipitation patterns, due to climate change, directly
influenced plant growth, impacting energy availability for herbivores and subsequently
carnivores. This underscores how changes in abiotic factors such as water availability can
drastically alter energy dynamics.
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Figure 8-B: Coral Reef Ecosystem Food Web (Rogers et al., 2015)
Implications for Engineering: Coral reef energy flow models are being used to develop
sustainable fisheries and marine protected areas. Understanding these energy dynamics can
also help in coral reef restoration projects, which aim to boost the productivity and resilience
of reefs.
A study in Northern Alaska revealed that energy transfer in the tundra is highly seasonal, with
peak energy flow occurring during the short summer when sunlight is most available (Begon
et al., 2006). During this time, plant productivity spikes, and herbivore populations increase,
providing a temporary energy boost to higher trophic levels. However, long winters
significantly reduce energy flow, resulting in lower biodiversity and simpler food chains
compared to other ecosystems.
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Figure 8-C: Tundra Ecosystem Food Web
Implications for Engineering: Knowledge of energy flow in the tundra has practical
applications for Arctic conservation efforts and sustainable development projects. It is
particularly relevant for designing infrastructure in these fragile ecosystems, as energy flow
and ecosystem productivity are highly sensitive to climate change and human disturbances.
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Figure 8-E: Energy Flow in Wetland Ecosystem
A study in the Florida Everglades showed that energy flow in wetlands is highly influenced
by water flow patterns and nutrient availability (Odum, 1971). When water levels dropped
due to drainage and water management projects, the energy available for herbivores and
decomposers decreased, leading to a decline in biodiversity and ecosystem health.
Implications for Engineering: Wetland ecosystems are key to water purification, flood
control, and carbon sequestration. Understanding energy flow in these systems allows
environmental engineers to design projects that restore natural water flow and support
ecosystem services.
A case study of urban ecosystems in New York City showed that energy flow is highly
modified by human infrastructure and consumption patterns (Pimm, 1982). Urban plants
provide habitat for insects and birds, while waste decomposition in landfills and composting
systems creates new pathways for energy recycling. Despite these interactions, energy flow is
less efficient compared to natural ecosystems, largely due to energy losses in transportation,
food supply chains, and waste management.
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Implications for Engineering: Understanding energy flow in urban ecosystems is critical for
designing more sustainable cities. Urban planners and environmental engineers are
increasingly focused on creating energy-efficient green spaces, promoting urban agriculture,
and enhancing waste management systems to reduce energy loss and improve ecosystem
health.
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10. Conclusion
Energy flow in ecosystems is a critical process that affects both natural and engineered
environments. The transfer of energy from producers to consumers and decomposers is
shaped by the laws of thermodynamics and is governed by both biological and physical
principles. From an engineering perspective, understanding these energy dynamics is
essential for designing sustainable ecosystems, mitigating environmental impacts, and
optimizing resource use. As human activities continue to impact ecosystems worldwide, the
need for innovative tools and approaches to measure and model energy flow will only grow
in importance.
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References
• Odum, E.P. (1971). Fundamentals of Ecology. Philadelphia: Saunders.
• Hall, C. A. S., & Moll, R. (1975). Methods of Assessing Aquatic Primary Productivity.
Primary Productivity of the Biosphere, 19–53.
• Gosz, J. R., Holmes, R. T., Likens, G. E., & Bormann, F. H. (1978). The flow of
energy in a forest ecosystem. Scientific American, 238(3), 92-103.
• Begon, M., Townsend, C.R., & Harper, J.L. (2006). Ecology: From Individuals to
Ecosystems. Blackwell Publishing.
• Fath, B. D., & Patten, B. C. (1999). "Review of the Foundations of Network Environ
Analysis." Ecosystems, 2(2), 167-179.
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