Osmoregulation & Excretion
Osmoregulation & Excretion
• osmolarity
◦ number of moles of solute per litre of solution
◦ isoosmotic, hyperosmotic, hypoosmotic
◦ response of animal cells - isotonic, hypertonic, hypotonic
• mechanisms in osmoregulation
◦ tolerance
‣ stenohaline - cannot tolerate substantial changes
‣ euryhaline - can survive large uctuations
◦ osmoconformer
‣ to be isoosmotic with its surroundings
‣ marine animals
• dehydrating environment
• mechanisms - active transport
◦ excretion of salt through kidneys and gills
◦ maintain high concentration of several other solutes - urea & TMAO
◦ osmoregulator
‣ to control internal osmolarity independent of that of the external environment
‣ freshwater animals - hyperosmotic
◦ in temporary waters
‣ enters a formant state - anhydrobiosis
• requires adaptations that keep cell
membranes intact
• trehalose - dissacharide
◦ protect the cells by replacing the water that is normally associated with proteins
and membrane lipids
◦ land animals
‣ reduce water loss
• waxy cuticle
• waxy layers of insect exoskeletons
• shells
• dead, keratinized skin cells
• nocturnal
◦ transport epithelia
‣ one or more layers of epithelial cells specialised for moving particular solutes in
controlled amounts in speci c directions
‣ adaptations
• tubular networks - surface area
• facing outside environment or by channels
• selective permeability & active transport
◦ active salt excretion
• excretion - removal of waste products
◦ ammonia
‣ toxic - ammonium can interfere with oxidative phosphrylation
‣ breakdown of nitrogenous molecules - proteins, amino acids
◦ excretory systems
‣ ltration
‣ reabsorption
‣ secretion
‣ excretion
• excretory tubules - nephrons & collecting
ducts
◦ atworms - protonephridia
‣ annelids e.g. earthworms
• metanephridia
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◦ concentrating urine - countercurrent multiplier systems
• adaptations
◦ % of cortical vs. juxtamedullary nephrons
◦ length of loops of Henle
◦ size of nephrons
• hormonal circuits
◦ kidney function, water balance, blood pressure
◦ ADH
‣ aquaporin