Chapt 19
Chapt 19
Biology
Sylvia S. Mader
Michael Windelspecht
Chapter 19
The Animals
BACTERIA
common
ARCHAEA
ancestor
(first cells)
Protists
Photosynthetic
protist
Plants
cell with nucleus
EUKARYA
Fungi
Heterotrophic
Protist Animals
common ancestor
Past Present
Time
Evolution of Animals 动物的进化
Multicellular eukaryotes. 多细胞真核生物。
• Like protists, plants, and fungi 像原生生物、植物和真菌
Chemoheterotrophs. 化学异养生物。
• Unlike autotrophic plants 与自养植物不同
• Like fungi, but animals ingest food and digest internally while fungi
digest food externally 像真菌一样,但动物摄入食物并在内部消化,而真菌在外部消化食物
Undergo sexual reproduction. 进行有性生殖。
• Begin life as a fertilized diploid egg 以受精二倍体卵开始生活
• Undergo developmental changes 经历发展变化
• Organism with specialized tissues 具有特殊组织的生物体
Possess nervous and muscular tissues 拥有神经和肌肉组织
• Allow for mobility (locomotion). 允许移动性(运动)。
The evolutionary tree of animals 动物的进化树
4. Coelom—body cavity 体腔 — 身体
Acoelomate – no body cavity 无体腔动物 — 没有身体腔隙腔隙
Pseudocoelomate - false body cavity 假体腔动物 - 假体腔
Coelomate – body cavity 体腔动物 — 身体腔隙
Figure 19.4 A modern look at the animal evolutionary tree.
Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
Lophotrochozoa Ecdysozoa
6
2. Symmetry 对称
Radial symmetry—animal is organized circularly 径向对称 - 动物是圆形组织的
Bilateral symmetry—animal has right and left halves 双侧对称——动物有右半部和
左半部
• Accompanied by cephalization—localization of brain and specialized
sensory organs 伴有头化——大脑和特殊感觉器官的定位
al
dors posterior
anterior
entr a l
v
a. b.
2. Protostomes and Deuterostomes 原口动物 vs. 后口动物
• Protostomes
Flatworms, roundworms, mollusks, annelids, arthropods 扁虫、蛔虫、软体动物、
环节动物、节肢动物
• Deuterostomes
Echinoderms and chordates 棘皮动物和脊索动物
ectoderm
mesoderm
endoderm
a. Acoelomate (flatworms)
b. Pseudocoelomate
(roundworms)
Coelomate—body cavity completely lined with
mesoderm 腔体——体腔完全衬有中胚层
• Mesentery (fold of membrane) supports internal organs
• In animals without a skeleton, the coelom acts as a hydrostatic
(fluid-filled) skeleton.
• 在没有骨骼的动物中,腔体充当流体静力(充满液体)的骨骼。肠系膜(膜褶皱)支持内脏器官
Phylogenetic Tree of Animals
Deuterostome
Common ancestor development Chordates
Deuterostomia
Bilateral Symmetry Echinoderms
3 tissue layers
body cavity
Arthropods
Ecdysozoa
Tissue
layers Roundworms
Bilateria
Annelids
Protostomia
Multicellularity Molluscs
Ancestor
Lophotrochozoa
Flatworms
Protostome
development Rotifers
Lophophores
Comb jellies
Radiata
Radial Symmetry Cnidarians
2 tissue layers
Sponges
No true tissues 11
Sponges 海绵
The Early Animals 早期动物
• Sponges: multicellular 海绵:多细胞
Aquatic, largely marine 水生,主要是海洋
Multicellular but lack organized tissues. No tissue layer 多细胞
但缺乏有组织的组织。无组织层
• Cellular level of organization 组织细胞级别
Filter feeder—filters food out of water using pores and collar
cell microvilli 滤食器—使用孔和领细胞微绒毛过滤食物中的水分
Reproduce both sexually and asexually 有性繁殖和无性繁殖
• Asexual fragmentation or budding 无性分裂或萌芽
Figure 19.8 Sponge Anatomy
Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
central sponge
cavity wall
amoeboid
cell
central pore
cavity porocyte
osculum epidermal
cell
spicule
Body wall
collar
flagellum nucleus
amoeboid cell
Yellow tube sponge Collar cell
© Andrew J. Martinez/Photo Researchers, Inc.
• Cnidarians: true tissues 刺胞动物:真正的组织
Radially symmetrical 径向对称
Aquatic, mostly marine 水生,主要是海洋
2 germ layers—ectoderm and endoderm2 胚层——外胚层和内胚
层
Tissue level of organization
• 组织层面的组织
Figure 19.9a, b Cnidarians.
tentacles
mouth
bud tentacles
a. Hydra
b. Sea anemone
Examples 例子
Planarians are free-living flatworms 涡虫是自由生活的扁虫
• Incomplete digestive 消化不完全
• Hermaphrodites 雌雄同体
Both male and female sex organs 男性和女性的性器官
Cross-fertilize 交叉施肥
Figure 19.10b Anatomy of a Planarian.
涡虫的解剖学。 Copyright © The McGraw-Hill Companies,
Inc. Permission required for reproduction or fluid
display. cilia
flame cell
ovary
transverse nerves
brain testis
sperm duct
gastrovascular
cavity pharynx
male
female
hooks
sucker
heart shell
coelom visceral mass
mantle
mantle
cavity
anus
gill
radula
mouth
a. foot
3 Classes of Mollusks: 3 类软体动物:
1. Gastropods1. 腹足类动物
• Examples: conchs, and snails 例如:海螺和蜗牛
• Herbivores or carnivores
• 食草动物或食肉动物
tentacle
gills
foot
Land snail
foot
mantle
Nudibranch
2. Cephalopods. 头足类动物
• Examples: octopuses, squids 例如:章鱼、鱿鱼
• Foot evolved into tentacles around head 脚进化成头
部周围的触手
• Well-developed nervous system and brain 发达的神
经系统和大脑
tentacles
eye
arm
eye
suckers
3. Bivalves 双壳类
• Examples: clams, oysters, scallops, mussels 例如:蛤蜊、牡蛎、
扇贝、贻贝
• 2 parts to the shell2 个零件到外壳
• Filter feeders 滤料喂料机
eye
tentacle valve
Scallop
valve
Mussels
• Annelids: segmented worms 环节动物:分节蠕虫
Segmented—externally in rings encircling body 分段 - 外部环绕主
体的环
Septa divide fluid-filled coelom 隔垫分流充满液体的腔体
• Used as hydrostatic skeleton 用作静压骨架
Complete digestive tract with specialized parts 具有特殊部件的完整消
化道
Nervous system has brain and ventral nerve cord 神经系统有脑神经
索和腹神经索
mouth pharynx
ventral
nerve cord brain
esophagus
ventral
blood vessel coelom
hearts
clitellum seminal
gizzard vesicle
nephridium crop
anus
dorsal
blood vessel
Annelids (3 groups)
1.Polychaetes 多毛纲
• Have many setae per segment. Example clam worm 每个体节有很多刚毛。例如:蛤蠕虫
2. Oligochaetes 寡毛纲
• Few setae per segment 每个体节有少量刚毛
• Example: Earthworm 例如:蚯蚓 Figure 19.
• Scavengers in soil 它们是土壤中的腐食者。 Examples of annelids.
spiraled
tentacles
setae
a. Clam worm
Figure 19.15c Examples of annelids.
anterior
sucker
posterior
sucker
No setae 没有刚毛
Most freshwater, but some marine or terrestrial 大多数生活在淡水中,但也有一些生活
在海洋或陆地上
Some free-living but most are fluid feeders 有些是自由生活的,但大多数是流食者
Hirudin is a powerful anticoagulant. 硫酸肝素是一种强力的抗凝血剂。
Roundworms 蛔虫
• Roundworms: Pseudocoelomates 蛔虫:假腔虫
Body cavity—pseudocoelom 体腔—假腔 Ascaris
Complete digestive tract 完整的消化道
• Both mouth and anus 口腔和肛门
Non-segmented 非分段
Occur almost everywhere 几乎无处不在
sea, fresh water, and soil 海洋、淡水和土壤
Free-living or parasitic 自由生活或寄生
Examples: parasites 示例:寄生虫
Ascaris (intestinal), 蛔虫(肠道),
Trichinosis (skeletal muscle)
旋毛虫病(骨骼肌)
Figure 19.16 Roundworm anatomy.
Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
mouth
pharynx
brain dorsal
nerve cord
excretory
pore cuticle
lateral a. Ascaris
ventral Nerve cord
nerve
cord
sperm
duct
gut
sperm pseudocoelom
muscle
layer
seminal
vesicle
muscle layer
testis
Trichinella larva
cyst
cloaca
spicules that
aid in sperm
transfer anus
b. Ascaris anatomy c. Trichinella ×124
2. Arachnids 蜘蛛动物
• Spiders 蜘蛛
• Scorpions 蝎子
• Mites 螨
• Centipedes 蜈蚣
• Millipedes 千足虫
• Arthropods
3. Insects Ants, Flies, Bees, 3. 昆虫蚂蚁、苍蝇、蜜蜂、
Termites, Mosquitoes, Butterflies, Roaches 白蚁、蚊子、蝴蝶、蟑螂
• Very numerous and diverse 数量众多,种类繁多
• Entomology—study of insects 昆虫学——昆虫研究
• Adapted to active life on land 适应陆地上的积极生活
Some have invaded aquatic habitats 有些已经入侵了水生栖息地
• Body—head, thorax, abdomen 身体——头部、胸部、腹部
• 3 pairs of legs, 1 or 2 pairs of wings or none
•
• 3 对腿, 1 或 2 对翅膀或没有
Phylogenetic Tree of Animals
Deuterostome
Common ancestor development Chordates
Deuterostomia
Bilateral Symmetry Echinoderms
3 tissue layers
body cavity
Arthropods
Ecdysozoa
Tissue
layers Roundworms
Bilateria
Annelids
Protostomia
Multicellularity Molluscs
Ancestor
Lophotrochozoa
Flatworms
Protostome
development Rotifers
Lophophores
Comb jellies
Radiata
Radial Symmetry Cnidarians
2 tissue layers
Sponges
No true tissues 33
The Deuterostomes 后口动物
• Deuterostomes: mouth forms second 嘴巴第二个形成
Echinoderms 棘皮动物
Larva is free-swimming with bilateral symmetry 幼虫自由游泳,双侧对称
Adults are radially symmetric no (head, brain) 成年体具有辐射对称,没有(头部、大脑
Locomotion depends on water vascular system 运动取决于水血管系统
No complex respiratory, excretory, or circulatory system 没有复杂的呼吸、排泄或循环
系统
4 characteristics:4 大特点:
1. Dorsal supporting rod called a notochord 背侧支撑
杆称为脊索
Vertebrates have an endoskeleton of cartilage or bone.
脊椎动物具有软骨或骨骼的内骨骼。
2. Dorsal tubular nerve cord 肾小管背侧神经索
3. Pharyngeal pouches 咽袋
4. Postanal tail 肛门后尾
Figure 19.25 The four chordate characteristics.
4 characteristics:
pharyngeal
pouches
dorsal tubular
nerve cord
notochord
postanal tail
• Evolutionary trends among the chordates
脊索动物的进化趋势
Vertebrates
Tetrapods
mammary glands
amniotic egg
*includes birds
limbs
lungs
bony skeleton
jaws
vertebrae
ancestral chordate
• Invertebrate chordates 无脊 Figure 19.26a
Invertebrate chordates.
椎动物脊索动物
Notochord never replaced by
vertebral column 脊索永不被脊椎柱所替代
Tunicates
• “Sea squirts” can squirt water from
siphon when disturbed 被囊状物覆盖的脊索动
物( Tunicates ) “海鞘”在受到干扰时可以从喷口
喷出水
Lancelets
• Marine chordates few cm long 剑尾动物
( Lancelets ) 海洋脊索动物,几厘米长
Vertebrate Chordates 脊椎动物脊索动物
Notochord replaced by vertebral column 脊索被脊柱取代
1.Fishes 鱼
Gills, 腮
Covered in scales, 覆盖着鳞片,
Reproduce by laying eggs, 通过产卵繁殖,
2-chambered heart 两腔心脏
1. Fishes
First vertebrates were jawless fishes 最早的脊椎动物
是无颌鱼类
• Examples: brain
nostril
• Perch, 例子:
• trout, 鲈鱼 scales
• salmon, 鳟鱼 mandible
gills
鲑鱼 kidney
gonad heart
• haddock 黑线鳕
intestine gallbladder liver
Feather anatomy
barbule
barb
shaft
nostril
ear opening
trachea
lung
testis esophagus
kidney
gizzard
vas deferens crop
ureter heart
liver
sternum
pancreas
rectumcloaca
Figure 19.33 Birds are well adapted for flight.
Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
5. Mammals 哺乳动物
Endotherms (like birds) 吸热
2 chief characteristics of
mammals 哺乳动物的 2 个主要特征
• Hair—provides insulation against
heat loss 头发—提供隔热材料,防止热量损失
• Mammary glands (milk glands)
enable female to feed young
without leaving them to find food
乳腺(乳腺)使雌性能够喂养幼崽,而无需离开它们寻找
食物
5. Mammals
Endotherms (like birds)
Three groups 三组
Monotremes 单体
Marsupials 有袋动物
Placental
胎盘的
Figure 19.35a
a. Monotremes 单体
Spiny anteater 多刺食蚁兽
Duckbill platypus 鸭嘴兽
• Have cloaca (like birds) and lay
hard-shelled amniote eggs 有泄殖腔(像
鸟一样)并产下硬壳羊膜蛋
Figure 19.35b, c.
b. Marsupials
b. 有袋动物
Mainly in Australia 主要在澳大利亚
• Born in immature condition 出生时未成熟
• Newborns complete development inside female’s pouch 新生儿在女性袋内完成
发育
• koalas, 考拉
• kangaroos, 袋鼠
• Tasmanian wolf (extinct) 塔斯马尼亚狼(已灭绝)
c. Placental Mammals 胎盘哺乳动物
• Vast majority of living mammals 绝大多数现存哺乳动物
• Extraembryonic membranes of amniote egg modified for internal
development of embryo within uterus of female 羊膜卵的胚外膜经过修饰,用于女性子宫
内胚胎的内部发育
• Adapted to active life on land 适应陆地上的积极生活
Well-developed brain—expansion of cerebral hemispheres 发达的大脑——大
脑半球的扩张
Four-chambered heart 四腔心脏
Internal temperature is constant (Endotherms) 内部温度恒定(吸热)
19.6 Human Evolution 人类进化
• All primates share a common ancestor. 所有灵长类动物都有一个共同的祖
先。
• Prosimians—lemurs, tarsiers, and lorises 普罗西米亚人——狐猴、眼
镜猴和懒猴
• Anthropoids—monkeys, apes, and humans 类人猿——猴子、猿类
和人类
Anthropoids
19.6 Human Evolution
• Evolutionary tree indicates humans most closely related to
African apes 进化树表明人类与非洲猿类关系最密切
Humans did not evolve from apes 人类不是从猿类进化而来的
Believed to share a common ancestor about 7 MYA
据信在大约 7 MYA 中有一个共同的祖先
human
chimpanzee
lemur baboon
Prosimians New World Monkeys Old World Monkeys Gibbons Apes and Chimpanzees Hominins
0
6
Hominid
12
18
Million Years Ago (MYA)
common ancestor
24 for apes and humans
30
36
42
48
common ancestor
54 for all primates
60
66
• Cro-Magnons (Homo sapiens) Cro-Magnons (智
人)
Modern appearance 现代外观
Advanced tools 高级工具
May have been first to make knifelike blades and
throw spears 可能是第一个制造刀状刀片和投掷长矛的人
Hunted cooperatively 合作狩猎
Perhaps first to have language 也许是第一个拥有语言的人
Culture included art 文化包括艺术