CasperJS is an open source navigation scripting and testing utility written in JavaScript that eases the process of defining navigation scenarios and provides functions for common tasks like filling forms, clicking links, taking screenshots, and testing websites. It uses the PhantomJS or SlimerJS headless browsers under the hood. The Casper API provides methods like start(), then(), run(), and exit() to control navigation and run test suites, as well as utilities like fill(), click(), evaluate(), and waitForSelector() to interact with pages.