Escarabajos de estiércol madre de dios
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1. This document describes a study of 60 species of dung beetles from the Amarakaeri Communal Reserve in Madre de Dios, Peru. 2. Images and descriptions are provided for each of the 60 identified species of dung beetles, which are classified into 12 genera. 3. The study was conducted by researchers from the Museo de Historia Natural de la Universidad Nacional San Antonio Abad del Cusco and the Smithsonian Conservation Biology Institute, with funding from the SCBI and Hunt Oil Exploration.
![Madre de Dios, PERÚ
Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve
1
Gorky Valencia1
, Alfonso Alonso2
, Rolando Coronel1
and Tremaine Gregory2
1
Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2
Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro.
Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell,
Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru.
(F) Female, (M) Male. Tick marks in images indicate 1 millimeter.
© All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016
1 Ateuchus connexus 2 Canthidium basipunctatum 3 Canthidium lentum 4 Canthidium onitoides
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
5 Canthon aberrans 6 Canthon aequinoctialis 7 Canthon femoralis 8 Canthon fulgidus
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
9 Coprophanaeus larseni (M) 10 Coprophanaeus telamon (F) 11 Coprophanaeus telamon (M) 12 Deltochilum amazonicum
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
13 Deltochilum carinatum 14 Deltochilum howdeni 15 Deltochilum larseni 16 Deltochilum orbiculare (F)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
17 Deltochilum orbiculare (M) 18 Deltochilum peruanum (F) 19 Deltochilum peruanum (M) 20 Dendropaemon attalus
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE](https://image.slidesharecdn.com/778perubeetlesofamarakaeri-1-160802143515/85/Escarabajos-de-estiercol-madre-de-dios-1-320.jpg)
![Madre de Dios, PERÚ
Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve
2
Gorky Valencia1
, Alfonso Alonso2
, Rolando Coronel1
and Tremaine Gregory2
1
Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2
Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro.
Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell,
Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru.
(F) Female, (M) Male. Tick marks in images indicate 1 millimeter.
© All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016
21 Dichotomius batesi 22 Dichotomius cf. fonsecae (M) 23 Dichotomius mamillatus (F) 24 Dichotomius mamillatus (M)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
25 Dichotomius ohausi (F) 26 Dichotomius ohausi (M) 27 Dichotomius prietoi 28 Dichotomius robustus
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
29 Dichotomius worontzowi 30 Eurysternus caribaeus 31 Eurysternus foedus 32 Eurysternus hypocrita
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
33 Eurysternus lanuginosus 34 Eurysternus nigrovirens 35 Eurysternus plebejus 36 Eurysternus vastiorum
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
37 Eurysternus wittmerorum 38 Ontherus alexis 39 Onthophagus haematopus (F) 40 Onthophagus haematopus (M)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE](https://image.slidesharecdn.com/778perubeetlesofamarakaeri-1-160802143515/85/Escarabajos-de-estiercol-madre-de-dios-2-320.jpg)
![Madre de Dios, PERÚ
Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve
3
Gorky Valencia1
, Alfonso Alonso2
, Rolando Coronel1
and Tremaine Gregory2
1
Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2
Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro.
Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell,
Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru.
(F) Female, (M) Male. Tick marks in images indicate 1 millimeter.
© All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016
41 Onthophagus onorei (F) 42 Onthophagus onorei (M) 43 Onthophagus osculatii (F) 44 Onthophagus osculatii (M)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
45 Onthophagus rubrescens (F) 46 Onthophagus xanthomerus (F) 47 Onthophagus xanthomerus (M) 48 Oxysternon conspicillatum (F)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
49 Oxysternon conspicillatum (M) 50 Oxysternon silenus (F) 51 Oxysternon silenus (M) 52 Oxysternon spiniferum (F)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
53 Oxysternon spiniferum (M) 54 Phanaeus bispinus (F) 55 Phanaeus bispinus (M) 56 Phanaeus cambeforti (F)
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
57 Phanaeus cambeforti (M) 58 Scybalocanthon aereus 59 Sylvicanthon bridarollii 60 Uroxys gorgon
SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE](https://image.slidesharecdn.com/778perubeetlesofamarakaeri-1-160802143515/85/Escarabajos-de-estiercol-madre-de-dios-3-320.jpg)
Escarabajos de estiércol madre de dios
- 1. Madre de Dios, PERÚ Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve 1 Gorky Valencia1 , Alfonso Alonso2 , Rolando Coronel1 and Tremaine Gregory2 1 Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2 Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro. Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell, Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru. (F) Female, (M) Male. Tick marks in images indicate 1 millimeter. © All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016 1 Ateuchus connexus 2 Canthidium basipunctatum 3 Canthidium lentum 4 Canthidium onitoides SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 5 Canthon aberrans 6 Canthon aequinoctialis 7 Canthon femoralis 8 Canthon fulgidus SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 9 Coprophanaeus larseni (M) 10 Coprophanaeus telamon (F) 11 Coprophanaeus telamon (M) 12 Deltochilum amazonicum SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 13 Deltochilum carinatum 14 Deltochilum howdeni 15 Deltochilum larseni 16 Deltochilum orbiculare (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 17 Deltochilum orbiculare (M) 18 Deltochilum peruanum (F) 19 Deltochilum peruanum (M) 20 Dendropaemon attalus SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
- 2. Madre de Dios, PERÚ Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve 2 Gorky Valencia1 , Alfonso Alonso2 , Rolando Coronel1 and Tremaine Gregory2 1 Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2 Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro. Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell, Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru. (F) Female, (M) Male. Tick marks in images indicate 1 millimeter. © All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016 21 Dichotomius batesi 22 Dichotomius cf. fonsecae (M) 23 Dichotomius mamillatus (F) 24 Dichotomius mamillatus (M) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 25 Dichotomius ohausi (F) 26 Dichotomius ohausi (M) 27 Dichotomius prietoi 28 Dichotomius robustus SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 29 Dichotomius worontzowi 30 Eurysternus caribaeus 31 Eurysternus foedus 32 Eurysternus hypocrita SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 33 Eurysternus lanuginosus 34 Eurysternus nigrovirens 35 Eurysternus plebejus 36 Eurysternus vastiorum SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 37 Eurysternus wittmerorum 38 Ontherus alexis 39 Onthophagus haematopus (F) 40 Onthophagus haematopus (M) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE
- 3. Madre de Dios, PERÚ Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve 3 Gorky Valencia1 , Alfonso Alonso2 , Rolando Coronel1 and Tremaine Gregory2 1 Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2 Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro. Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell, Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru. (F) Female, (M) Male. Tick marks in images indicate 1 millimeter. © All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016 41 Onthophagus onorei (F) 42 Onthophagus onorei (M) 43 Onthophagus osculatii (F) 44 Onthophagus osculatii (M) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 45 Onthophagus rubrescens (F) 46 Onthophagus xanthomerus (F) 47 Onthophagus xanthomerus (M) 48 Oxysternon conspicillatum (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 49 Oxysternon conspicillatum (M) 50 Oxysternon silenus (F) 51 Oxysternon silenus (M) 52 Oxysternon spiniferum (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 53 Oxysternon spiniferum (M) 54 Phanaeus bispinus (F) 55 Phanaeus bispinus (M) 56 Phanaeus cambeforti (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE 57 Phanaeus cambeforti (M) 58 Scybalocanthon aereus 59 Sylvicanthon bridarollii 60 Uroxys gorgon SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE